Psephophorus
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''Psephophorus'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
sea turtle Sea turtles (superfamily Chelonioidea), sometimes called marine turtles, are reptiles of the order Testudines and of the suborder Cryptodira. The seven existing species of sea turtles are the flatback, green, hawksbill, leatherback, loggerhe ...
that lived from the Oligocene to the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, North America, and New Zealand. It was first named by
Hermann von Meyer Christian Erich Hermann von Meyer (3 September 1801 – 2 April 1869), known as Hermann von Meyer, was a German palaeontologist. He was awarded the 1858 Wollaston medal by the Geological Society of London. Life He was born at Frankfurt am Ma ...
in 1847, and contains seven species, ''P. polygonus'', ''P. calvertensis'', ''P. eocaenus'', ''P. oregonesis'', ''P. californiensis'', ''P. rupeliensis'',''Proceedings''; page 8. By the Zoological Society of London; published 1891. Retrieved on June 28th, 2008. ''P. scaldii'', and a species discovered in 1995, ''P. terrypratchetti''. ''Psephophorus'' is the only
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
dermochelyid turtle found in Europe. One species of ''Psephophorus'' could measure up to ten feet in length.


Discovery and identification

Von Meyer originally named ''Psephophorus'' in 1846. At first he was unable to identify the creature beyond its
dermal plate The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided in ...
s, but when he later received a drawing he was able to describe the specimen, which was then in Pressburg, as a fragment of a carapace, which contained seventy
bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
s. In 1879, H. G. Seeley was asked to study the ''Psephophorus'' specimen by
Franz Ritter von Hauer Franz Ritter von Hauer, or Franz von Hauer (January 30, 1822 – March 20, 1899), was an Austrian geologist. Biography Hauer was born in Vienna, the son of Joseph von Hauer (1778-1863), who was equally distinguished as a high Austrian offic ...
, the Director of the Austro-Hungarian
Imperial and Royal Geological Survey Imperial is that which relates to an empire, emperor, or imperialism. Imperial or The Imperial may also refer to: Places United States * Imperial, California * Imperial, Missouri * Imperial, Nebraska * Imperial, Pennsylvania * Imperial, Te ...
. Up until then, the specimen's identity had been undeterminable, with even Seeley describing it at first to seem like "the dermal covering of an Edentate closely allied to the Armadilloes." Seeley examined some bone fragments and concluded the specimen was that of a reptilian creature, furthermore a chelonid. It also proved to be more closely related to '' Sphargis'' than any other type in the Chelonian order.


Relation to modern Leatherback sea turtles

For a long time, modern
Leatherback The leatherback sea turtle (''Dermochelys coriacea''), sometimes called the lute turtle or leathery turtle or simply the luth, is the largest of all living turtles and the heaviest non-crocodilian reptile, reaching lengths of up to and weights ...
sea turtles (''Dermochelys'') were believed to be descended directly from ''Psephophorus'', specifically the species ''P. polygonus''. However, a 1996 analysis by Wood et al. proved that most of the
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
in the two genera were not connected, meaning ''Psephophorus'' could not be a direct ancestor of the modern leatherbacks. The platelets on ''Psephophorus'' are quite similar to those on '' Dermochelys'', despite differences in outer
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
and size. The platelet comprises an external compact layer and an internal zone of
cancellous bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, and ...
.


Species

*''Psephophorus polygonus'' is the type
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, and was discovered by von Meyer in 1846. Fossils of the species have shown bony, carapacial ridges, adjacent, small and polygonal bony ossicles which measure about 22 × 25 mm. When the ossicles extend further along the longitudinal axis, their size increases up to an average of 33 × 41 mm. *''Psephophorus calvertensis'' was first named by Palmer in 1909. It was so named for being found in the
Calvert Formation The Calvert Formation is a geologic formation in Maryland, Virginia, and Delaware. It preserves fossils dating back to the early to middle Miocene epoch of the Neogene period. The formation is a destination for amateur fossil hunters as well as ...
. ''P. calvertensis'' is a rarely found specimen. *''Psephophorus eocaenus'' was first named by Andrews in 1901. *''Psephophorus californiensis'' was first named by Gilmore in 1937. *''Psephophorus terrypratchetti'' was discovered in the 1990s by Richard Köhler in New Zealand and named in 1995 after the author
Terry Pratchett Sir Terence David John Pratchett (28 April 1948 – 12 March 2015) was an English humourist, satirist, and author of fantasy novels, especially comical works. He is best known for his '' Discworld'' series of 41 novels. Pratchett's first no ...
, who wrote a series of
fantasy books Fantasy literature is literature set in an imaginary universe, often but not always without any locations, events, or people from the real world. Magic, the supernatural and magical creatures are common in many of these imaginary worlds. Fan ...
set on a world carried on the back of a giant turtle. This species reached in body length which is more than 80% bigger than ''P. eocaenus''.


See also

*
List of organisms named after famous people (born 1900–1949) In biological nomenclature, organisms often receive scientific names that honor a person. A taxon (e.g. species or genus; plural: taxa) named in honor of another entity is an eponymous taxon, and names specifically honoring a person or persons ar ...
* '' Archelon'' * '' Gigantatypus'' *
Largest prehistoric animals The largest prehistoric animals include both vertebrate and invertebrate species. Many of them are described below, along with their typical range of size (for the general dates of extinction, see the link to each). Many species mentioned might ...


References


Further reading

* * * *


External links

* * * * * * {{Taxonbar, from=Q7254102 Dermochelyidae Fossil taxa described in 1995 Terry Pratchett Oligocene turtles Miocene turtles Pliocene turtles Cenozoic reptiles of Europe Cenozoic reptiles of North America Oligocene genus first appearances Pliocene genus extinctions Prehistoric turtle genera Taxa named by Christian Erich Hermann von Meyer Extinct turtles