A psammosere is a

seral community A seral community is an intermediate stage found in ecological succession in an ecosystem advancing towards its climax community. In many cases more than one seral stage evolves until climax conditions are attained. A prisere is a collection of se ...

, an

ecological succession Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades (for example, after a wildfire) or more or less. Bacteria allows for the cycling of nutrients such as ca ...

that began life on newly exposed coastal sand. Most common psammoseres are sand

dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, f ...

systems. In a psammosere, the

organism In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and ...

s closest to the sea will be

pioneer species Pioneer species are hardy species that are the first to colonize barren environments or previously biodiverse steady-state ecosystems that have been disrupted, such as by wildfire. Pioneer flora Some lichens grow on rocks without soil, so m ...

: salt-tolerant species such as

littoral The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal areas ...

algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...

and

glasswort The glassworts are various succulent, annual halophytic plants, that is, plants that thrive in saline environments, such as seacoasts and salt marshes. The original English glasswort plants belong to the genus ''Salicornia'', but today the glass ...

with

marram grass ''Ammophila'' (synonymous with ''Psamma'' P. Beauv.) is a genus of flowering plants consisting of two or three very similar species of grasses. The common names for these grasses include marram grass, bent grass, and beachgrass. These grasses ar ...

stabilising the dunes. Progressing inland many characteristic features change and help determine the natural succession of the dunes. For instance, the drainage slows down as the land becomes more compact and has better soils, and the pH drops as the proportion of seashell fragments reduces and the amount of

humus In classical soil science, humus is the dark organic matter in soil that is formed by the decomposition of plant and animal matter. It is a kind of soil organic matter. It is rich in nutrients and retains moisture in the soil. Humus is the Lati ...

increases. Sea purslane,

sea lavender ''Limonium'' is a genus of 120 flowering plant species. Members are also known as sea-lavender, statice, caspia or marsh-rosemary. Despite their common names, species are not related to the lavenders or to rosemary. They are instead in Plumbag ...

meadow grass ''Poa'' is a genus of about 570 species of grasses, native to the temperate regions of both hemispheres. Common names include meadow-grass (mainly in Europe and Asia), bluegrass (mainly in North America), tussock (some New Zealand species), a ...

and heather eventually grade into a typical non-maritime terrestrial eco-system. The first trees (or pioneer trees) that appear are typically fast-growing trees such as

birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...

willow Willows, also called sallows and osiers, from the genus ''Salix'', comprise around 400 speciesMabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge. of typically deciduous trees and shrubs, found primarily on moist s ...

rowan The rowans ( or ) or mountain-ashes are shrubs or trees in the genus ''Sorbus ''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' (''s.l.'') are commonly known as whitebeam, r ...

. In turn these will be replaced by slow-growing, larger trees such as

ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non- gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...

and

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

. This is the

climax community In scientific ecology, climax community or climatic climax community is a historic term for a community of plants, animals, and fungi which, through the process of ecological succession in the development of vegetation in an area over time, hav ...

, defined as the point where a plant succession does not develop any further because it has reached equilibrium with the

environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...

, in particular the

climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologic ...

. In an idealised coastal psammosere model, at the seaward edge of the sand dune the pH of the soil is typically alkaline/neutral with a pH of 7.0/8.0 particularly where shell fragments provide a significant component of the sand. Tracking inland across the dunes a

podsol In soil science, podzols are the typical soils of coniferous or boreal forests and also the typical soils of eucalypt forests and heathlands in southern Australia. In Western Europe, podzols develop on heathland, which is often a construct of huma ...

develops with a pH of 5.0/ 4.0 followed by mature podsols at the climax with a pH of 3.5 - 4.5.

External links

Case study of a psammosere
{{Ecology-stub Ecological succession

See also

External links