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''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
s and rodents which they actively hunt. All species in the genus are venomous, and the venom is considered mild and not dangerous to humans.


Etymology

The generic name ''Psammophis'' was coined by the Austrian herpetologist
Leopold Fitzinger Leopold Joseph Franz Johann Fitzinger (13 April 1802 – 20 September 1884) was an Austrian zoologist. Fitzinger was born in Vienna and studied botany at the University of Vienna under Nikolaus Joseph von Jacquin. He worked at the Vienna Naturhis ...
in 1826, a compound of the Hellenistic Greek ψαμμο, "sand" + Classical Greek ὄϕις, "snake", thus sand-snake.


Description

In the genus ''Psammophis'' the maxillary teeth are 10 to 13 in number, with one or two in the middle much enlarged and fang-like, preceded and followed by an interspace; the two posterior teeth are grooved. The anterior mandibular teeth are long, and the posterior teeth are small. The head is elongated and distinct from the neck, with an angular canthus rostralis. The eye is rather large, with a round pupil. The body is elongated and cylindrical with smooth dorsal scales in 15 or 17 rows at midbody, with apical pits. The ventral scales are rounded or obtusely angulate laterally, and the tail is long with the subcaudals in two rows.


Geographic range

Thirty-three species of ''Psammophis'' are known, from Africa and southern Asia. One fossil species, ''
Psammophis odysseus ''Psammophis odysseus'' is an extinct species of terrestrial snake belonging to the family Psammophiidae. It lived around 5.5 million years ago and was discovered in Salobreña, Spain by Georgios Georgalis. It is believed to have inhabited A ...
'', is known to have inhabited the Iberian Peninsula during the Miocene, indicating that the genus managed to successfully colonize mainland Europe in the wake of the Messinian salinity crisis, only to be extirpated from there shortly afterwards.


Behavior

''Psammophis'' species are chiefly sand-snakes, but they are also found on low bushes.


Diet

Species in the genus ''Psammophis'' feed principally on lizards. Boulenger GA (1890). ''The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia.'' London: Secretary of State for India in Council. (Taylor and Francis, printers). xviii + 541 pp. (Genus ''Psammophis'', p. 365, Figure 107).


Reproduction

All species in the genus ''Psammophis'' are
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
. Branch, Bill (2004). ''Field Guide to Snakes and other Reptiles of Southern Africa''. Third Revised edition, Second impression. Sanibel Island, Florida: Ralph Curtis Books. 399 pp. . (Genus ''Psammophis'', p. 89).


Species

The following 35 extant species are recognized as being valid: *''
Psammophis aegyptius ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. A ...
'' *''
Psammophis afroccidentalis ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. A ...
'' *''
Psammophis angolensis ''Psammophis angolensis'', sometimes known as the dwarf sand snake, is a species of snake in the family Psammophiidae reaching a maximum length of 50 cm, but averaging 30 cm. The snake actively forages for lizards and frogs.MARAIS, J. 2 ...
'' *''
Psammophis ansorgii ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. A ...
'' *''
Psammophis biseriatus ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. A ...
'' *'' Psammophis brevirostris'' *'' Psammophis condanarus'' *''
Psammophis crucifer ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. ...
'' *''
Psammophis elegans ''Psammophis elegans'', the elegant sand racer, is a species of psammophiid snake. It is found in tropical Africa. See also * List of reptiles of Ghana References Psammophis Reptiles described in 1802 {{snake-stub ...
'' *''
Psammophis indochinensis ''Psammophis indochinensis'', also known as the Indo-Chinese sand snake, is a species of snake in the family Psammophiidae. Its conservation status is of "least concern". It is found at low elevations in Thailand, Cambodia, Laos, and Myanma ...
'' *'' Psammophis jallae'' *''
Psammophis leightoni ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. ...
'' *''
Psammophis leithii ''Psammophis leithii'', commonly called the Pakistan sand racer, Pakistani ribbon snake, or Leith's sand snake, is a species of rear-fanged snake in the family Psammophiidae. The species is native to South Asia. It is harmless to humans. Etymol ...
'' *'' Psammophis leopardinus'' *''
Psammophis lineatus ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively ...
'' *''
Psammophis lineolatus ''Psammophis lineolatus'', commonly known as steppe ribbon racer or arrow snake, is a species of Mildly venomous snake (Not Harmful for Humans) in the family Lamprophiidae. It is located in northern and central Asia, from north western China, M ...
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Psammophis longifrons ''Psammophis longifrons'', the stout sand snake or long sand racer, is a species of snake found in India. It can grow to a maximum length of . Description Greyish above in front, browner behind, the scales edged with black, particularly those o ...
'' *'' Psammophis mossambicus'' *'' Psammophis notostictus'' *''
Psammophis orientalis ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively ...
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Psammophis phillipsii ''Psammophis phillipsii'', also known by its common name olive grass racer, is a species from the genus ''Psammophis ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and ...
'' *''
Psammophis praeornatus ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. A ...
'' *'' Psammophis pulcher'' *''
Psammophis punctulatus ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. ...
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Psammophis rukwae ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. ...
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Psammophis schokari The Schokari sand racer (''Psammophis schokari'') is a species of Psammophiidae, psammophiid snake found in parts of Asia and Africa. ''Psammophis schokari aegyptius'' has been elevated to species status. Many people refer to snakes in the genus ...
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Psammophis sibilans ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. A ...
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Psammophis subtaeniatus ''Psammophis subtaeniatus'', the western yellow-bellied sand snake, is a snake found in northern Southern Africa; more specifically the north of KwaZulu-Natal and further north to Mozambique, Zimbabwe, Gauteng, North West, Limpopo, and Eswat ...
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Psammophis sudanensis ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. A ...
'' *''
Psammophis tanganicus ''Psammophis'' is a genus of snakes in the Family (biology), family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which the ...
'' *'' Psammophis trigrammus'' *''
Psammophis turpanensis ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively hunt. ...
'' *''
Psammophis zambiensis ''Psammophis'' is a genus of snakes in the family Psammophiidae. The genus comprises 33 species, which are found in Africa and Asia.. www.reptile-database.org. ''Psammophis'' are diurnal and prey on lizards and rodents which they actively ...
''


Fossil species

*†''
Psammophis odysseus ''Psammophis odysseus'' is an extinct species of terrestrial snake belonging to the family Psammophiidae. It lived around 5.5 million years ago and was discovered in Salobreña, Spain by Georgios Georgalis. It is believed to have inhabited A ...
'' '' Nota bene'': A
binomial authority In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
in parentheses indicates that the species was originally described in a genus other than ''Psammophis''.


References


Further reading

* Fitzinger LI (1826). ''Neue Classification der Reptilien nach ihre natürlichen Verwandtschaften. Nebst einer Verwandtshafts-tafel und einem Verzeichnisse der Reptilien-Sammlung des K.K. zoologischen Museum's zu Wien''. Vienna: J.G. Heubner. five unnumbered + 67 pp. + one plate. (''Psammophis'', new genus, p. 29). (in German and Latin). * {{Authority control Psammophis Taxa named by Leopold Fitzinger Snake genera