Protobothrops Kaulbacki
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''Protobothrops kaulbacki'', commonly known as the Kaulback's lance-headed pitviper or Kaulback's lance-headed pit viper,Gumprecht A, Tillack F, Orlov NL,
Captain A Captain is a title, an appellative for the commanding officer of a military unit; the supreme leader of a navy ship, merchant ship, aeroplane, spacecraft, or other vessel; or the commander of a port, fire or police department, election precinct, e ...
, Ryabov S (2004). ''Asian Pit vipers''. Berlin: Geitje Books. First Edition. 368 pp. .
is a
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . crotaline snakes (from grc, κρόταλον ''krotalon'' castanet), or pit adders, are a subfa ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
. No
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are currently recognized.


Etymology

The specific name, ''kaulbacki'', is in honor of British explorer
Ronald Kaulback Ronald John Henry Kaulback (23 July 1909 – 2 October 1995) was a British explorer, botanist and geographer. He was the son of Colonel H. A. Kaulback and had a younger brother Roy (known as Bill). After studying at the University of Cambridge and ...
.


Description

Adult males of ''P. kaulbacki'' may attain a total length of , which includes a tail long. Females may grow longer: maximum total length , tail .
Dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
, it is green, with a vertebral series of dark angular spots, which may be joined to form a zigzag stripe. The top of the head is black with yellow stripes. Ventrally, except for the whitish throat area, it is gray with large squarish or crescent-shaped yellow spots. Smith MA (1943). ''The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. III.—Serpentes''. London: Secretary of State for India. (Taylor and Francis, printers). xii + 583 pp. (''Protobothrops kaulbacki'', p. 512). Scalation includes 25 rows of
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
at midbody, 201–212
ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that conta ...
, 66–78
subcaudal scales In snakes, the subcaudal scales are the enlarged plates on the underside of the tail.Wright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . These scales may be either single or divided (pair ...
, and 8
supralabial scales In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
of which the third is the largest.


Geographic range

''Protobothrops kaulbacki'' had originally been known only from the type locality, which is "Pangnamdim, north of the Triangle,
Upper Burma Upper Myanmar ( my, အထက်မြန်မာပြည်, also called Upper Burma) is a geographic region of Myanmar, traditionally encompassing Mandalay and its periphery (modern Mandalay, Sagaing, Magway Regions), or more broadly speak ...
" (
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
). In 2005 it was reported also from
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
(China). Rao D, Zhao E (2005). "A New Record from China – ''Protobothrops kaulbacki'' (Reptilia, Serpentes, Viperidae)". ''Acta Zootaxonomica Sinica'' 30: 209-211. It also occurs in
Arunachal Pradesh Arunachal Pradesh (, ) is a state in Northeastern India. It was formed from the erstwhile North-East Frontier Agency (NEFA) region, and became a state on 20 February 1987. It borders the states of Assam and Nagaland to the south. It shares int ...
(NE India).


Reproduction

''Protobothrops kaulbacki'' is
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
. The adult female lays a
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
of 6–32
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s in a hole in the ground, and then remains with the eggs to guard them. Eggs measure 48–53 mm x 26–27 mm (about 2 in x 1 in). Each hatchling is 26–27 cm (about 10½ inches) in total length.


References


Further reading

*Kraus, Fred; Mink, Daniel G.; Brown, Wesley M. (1996). "Crotaline Intergeneric Relationships Based on Mitochondrial DNA Sequence Data". ''Copeia'' 1996 (4): 763–773. (''Protobothrops kaulbacki'', new combination). * Smith MA (1940). "The Amphibians and Reptiles obtained by Mr. Ronald Kaulback in Upper Burma". ''Records of the Indian Museum'' 42: 465–486. (''Protobothrops kaulbacki'', new species, p. 485, Plate VIII, Figure 5). kaulbacki Snakes of China Reptiles of India Reptiles of Myanmar Fauna of Tibet Reptiles described in 1940 Taxa named by Malcolm Arthur Smith {{Snake-stub