Protoalligator Huiningensis
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''Protoalligator'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
alligatoroid Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more c ...
crocodilian Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...
found in the
Anhui Anhui , (; formerly romanized as Anhwei) is a landlocked province of the People's Republic of China, part of the East China region. Its provincial capital and largest city is Hefei. The province is located across the basins of the Yangtze River ...
province of China and lived during the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
.


Taxonomy

''P. huiningensis'', was originally named ''Eoalligator huiningensis'', which therefore remains the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of the genus.Young CC. 1982. A Cenozoic crocodile from Huaining, Anhui. Selected Works of Yang Zhongjian. China: Academia Sinica. 47-48 But then a 2016 study of Chinese alligators by Wang ''et al.'' found the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
'' Eoalligator chunyii'' to be a junior
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
of ''
Asiatosuchus nanlingensis ''Asiatosuchus'' is an extinct genus of crocodyloid crocodilians that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to ''Asiatosuchus'' since the genus was named in 1940. These specie ...
'' and a basal member of
Crocodylidae Crocodiles (family Crocodylidae) or true crocodiles are large semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term crocodile is sometimes used even more loosely to include all extant mem ...
. However, a subsequent study by Wu ''et al.'' disagreed with the synonymy of ''"Asiatosuchus" nanlingensis'' and ''Eoalligator chunyii'', finding them to be distinct based on first-hand studies of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
s and cladistic analysis, although they agreed in classifying ''E. chunyii'' as a crocodyloid. In 2016, Wang ''et al.'' also found ''Eoalligator huiningensis'' to be an
alligatoroid Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more c ...
distinct from ''E. chunyii'', and thus moved ''Eoalligator huiningensis'' to the new genus ''Protoalligator''. A 2019 study by Massonne ''et al.'' included additional taxa from
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
and found ''
Eoalligator ''Eoalligator'' is an extinct genus of alligatoroid crocodilian from Paleocene deposits in China. Systematics ''Eoalligator'' was originally classified as an extinct relative of alligators. The holotype of ''E. chunyii'', IVPP V2716, is a partia ...
'' and ''Protoalligator'' to be related and as basal members of Alligatoroidea, as shown in the cladogram below:


References

Crocodilians Paleocene reptiles of Asia Paleocene crocodylomorphs Fossil taxa described in 2016 Prehistoric pseudosuchian genera {{paleo-archosaur-stub