Proteoteras Aesculana
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''Proteoteras aesculana'', the maple twig borer, early proteoteras or maple tip moth, is a
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
of the family Tortricidae. It is found from coast to coast in the northern United States, south to
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Miss ...
in the east and to
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
in the west. It has a scattered distribution in
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, from
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native Eng ...
to southern
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
. The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
is 11–18 mm. Adults are on wing from April to October in the northern United States. In California, adults have been collected as early as February and as late as September, suggesting that there are multiple generations in the
San Francisco Bay Area The San Francisco Bay Area, often referred to as simply the Bay Area, is a populous region surrounding the San Francisco, San Pablo, and Suisun Bay estuaries in Northern California. The Bay Area is defined by the Association of Bay Area Go ...
. There is one generation per year in the eastern part of the range. Larvae have been recorded on
boxelder ''Acer negundo'', the box elder, boxelder maple, Manitoba maple or ash-leaved maple, is a species of maple native to North America. It is a fast-growing, short-lived tree with opposite, compound leaves. It is sometimes considered a weedy or inv ...
, silver maple, sugar maple and bigleaf maple, but probably also feed on other maple species. The larvae hollow out dormant buds and seeds in fall and continue to feed on dormant buds in spring. During the growing season, larvae bore in fresh shoots, often destroying them and preventing terminal growth. The larvae mix
frass Frass refers loosely to the more or less solid excreta of insects, and to certain other related matter. Definition and etymology ''Frass'' is an informal term and accordingly it is variously used and variously defined. It is derived from the ...
(which is ejected from the galleries) with webbing to form shelters around the entrances.


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ImagesBug Guide
Eucosmini Moths of North America Moths described in 1881 {{Eucosmini-stub