Procellaria Aequinoctialis
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The white-chinned petrel (''Procellaria aequinoctialis'') also known as the Cape hen and shoemaker, is a large
shearwater Shearwaters are medium-sized long-winged seabirds in the petrel family Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwa ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which also ...
. It ranges around the
Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-small ...
as far north as southern Australia, Peru and Namibia, and breeds colonially on scattered islands. The white-chinned petrel was formerly considered to be
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organ ...
with the
spectacled petrel The spectacled petrel (''Procellaria conspicillata'') is a rare seabird that nests only on the high western plateau of Inaccessible Island in the South Atlantic Tristan da Cunha group. It is one of the largest petrels that nests in burrows. This ...
(''Procellaria conspicillata'').


Taxonomy

In 1747 the English naturalist George Edwards included an illustration and a description of the white-chinned petrel in the second volume of his ''A Natural History of Uncommon Birds''. He used the English name "The great Black Peteril" and based his hand-coloured etching on a preserved specimen that had been brought to London. He believed that it had been collected near the
Cape of Good Hope The Cape of Good Hope ( af, Kaap die Goeie Hoop ) ;''Kaap'' in isolation: pt, Cabo da Boa Esperança is a rocky headland on the Atlantic coast of the Cape Peninsula in South Africa. A common misconception is that the Cape of Good Hope is t ...
. When in 1758 the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' for the tenth edition, he placed the white-chinned petrel with the other petrels in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Procellaria Procellaria is a genus of Southern Ocean long-winged seabirds related to prions, and within the order Procellariiformes. The black petrel (''Procellaria parkinsoni'') ranges in the Pacific Ocean, and as far north as Central America. The spect ...
''. Linnaeus included a brief description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Procellaria aequinoctialis'' and cited Edwards' work. There are two subspecies: * ''P. a. aequinoctialis'' - breeds on islets of the temperate and subantarctic
South Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
and
South Indian South India, also known as Dakshina Bharata or Peninsular India, consists of the peninsular southern part of India. It encompasses the Indian states of Andhra Pradesh, Karnataka, Kerala, Tamil Nadu, and Telangana, as well as the union territo ...
Oceans, winters throughout them * ''P. a. steadi'' - breeds on the subantarctic islands of
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
, winters throughout the South Pacific Ocean Both subspecies were formerly considered synonymous with each other, but were recognized as distinct by the
International Ornithological Congress International is an adjective (also used as a noun) meaning "between nations". International may also refer to: Music Albums * ''International'' (Kevin Michael album), 2011 * ''International'' (New Order album), 2002 * ''International'' (The T ...
in 2022 based on strong genetic distinctiveness. The genus name ''Procellaria'' is from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''procella'' meaning "storm" or "gale. The specific epithet ''aequinoctialis'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
meaning ''equinoctial'' (''aequinoctium'' is the Latin word for "equinox"). The word ''petrel'' is derived from
Saint Peter Saint Peter; he, שמעון בר יונה, Šimʿōn bar Yōnāh; ar, سِمعَان بُطرُس, translit=Simʿa̅n Buṭrus; grc-gre, Πέτρος, Petros; cop, Ⲡⲉⲧⲣⲟⲥ, Petros; lat, Petrus; ar, شمعون الصفـا, Sham'un ...
and the story of his walking on water, referring to their appearing to run on the water's surface when taking to the air.Gotch, A. T. (1995) Other English names for this species are Cape hen and shoemaker. The white-chinned petrel is a member of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Procellariidae The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which also ...
, and the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Procellariiformes Procellariiformes is an order of seabirds that comprises four families: the albatrosses, the petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still called tubenoses in English, procellariiforms are of ...
. As a member of the Procellariiformes, they share certain identifying features. First, they have nasal passages that attach to the upper bill called
naricorns A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
. Although the nostrils on the petrel are on top of the upper bill. The bills of Procellariiformes are also unique in that they are split into between seven and nine horny plates. They produce a
stomach oil Stomach oil is the light oil composed of neutral dietary lipids found in the proventriculus (fore-gut) of birds in the order Procellariiformes. All albatrosses, procellarids (gadfly petrels and shearwaters) and northern and austral storm petrel ...
made up of
wax ester A wax ester (WE) is an ester of a fatty acid and a fatty alcohol. Wax esters comprise the main components of three commercially important waxes: carnauba wax, candelilla wax, and beeswax.. Wax esters are formed by combining one fatty acid with o ...
s and
triglycerides A triglyceride (TG, triacylglycerol, TAG, or triacylglyceride) is an ester derived from glycerol and three fatty acids (from ''tri-'' and ''glyceride''). Triglycerides are the main constituents of body fat in humans and other vertebrates, as we ...
that is stored in the
proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects. Birds The ...
. This can be sprayed out of their mouths as a defence against predators and as an energy rich food source for chicks and for the adults during their long flights. Finally, they also have a
salt gland The salt gland is an organ for excreting excess salts. It is found in the cartilaginous fishes subclass elasmobranchii (sharks, rays, and skates), seabirds, and some reptiles. Salt glands can be found in the rectum of sharks. Birds and reptile ...
that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a high saline solution from their nose.


Description

The white-chinned petrel measures in length and has a wingspan of . Males are heavier and average whereas females weigh around . Not only is it the largest ''Procellaria'' petrel but is also the largest species in its
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
outside of the
giant petrel Giant petrels form a genus, ''Macronectes'', from the family Procellariidae, which consists of two species. They are the largest birds of this family. Both species are restricted to the Southern Hemisphere, and though their distributions overlap ...
s. This large petrel is sooty-black and has some white on its throat and chin, more so in the Indian Ocean sector than the Atlantic. Its
primaries Primary elections, or direct primary are a voting process by which voters can indicate their preference for their party's candidate, or a candidate in general, in an upcoming general election, local election, or by-election. Depending on the c ...
can have a silvery appearance underneath. Its
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
may be horn or yellow with some black between the nostrils. It has black legs and feet. When it flies, it mixes slow wing beats with glides. Although normally quiet, it will rattle or groan while at its colony.


Distribution and habitat

The white-chinned petrel utilises many islands during the breeding season. 2,000,000 pairs breed on
South Georgia South Georgia ( es, Isla San Pedro) is an island in the South Atlantic Ocean that is part of the British Overseas Territory of South Georgia and the South Sandwich Islands. It lies around east of the Falkland Islands. Stretching in the east ...
, Between 175,000 and 226,000 pairs are on the
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the sub-Antarctic constituting one of the two exposed parts of the Kerguelen Plateau, a large ...
, and 100,000 pairs on
Disappointment Island Disappointment Island is one of seven uninhabited islands in the Auckland Islands archipelago, in New Zealand. It is south of the country's main South Island and from the northwest end of Auckland Island. It is home to a large colony of white ...
. Some also breed on the
Crozet Islands The Crozet Islands (french: Îles Crozet; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarcti ...
,
Prince Edward Islands The Prince Edward Islands are two small uninhabited islands in the sub-Antarctic Indian Ocean that are part of South Africa. The islands are named Marion Island (named after Marc-Joseph Marion du Fresne, 1724–1772) and Prince Edward Island ...
,
Campbell Islands The Campbell Islands (or Campbell Island Group) are a group of subantarctic islands, belonging to New Zealand. They lie about 600 km south of Stewart Island. The islands have a total area of , consisting of one big island, Campbell Isla ...
,
Auckland Islands The Auckland Islands (Māori: ''Motu Maha'' "Many islands" or ''Maungahuka'' "Snowy mountains") are an archipelago of New Zealand, lying south of the South Island. The main Auckland Island, occupying , is surrounded by smaller Adams Island, ...
,
Antipodes Islands The Antipodes Islands ( Maōri: Moutere Mahue; "Abandoned island") are inhospitable and uninhabited volcanic islands in subantarctic waters to the south of – and territorially part of – New Zealand. The 21 km2 archipelago lies 860  ...
, and the
Falkland Islands The Falkland Islands (; es, Islas Malvinas, link=no ) is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and about from Cape Dubouzet ...
. During the non-breeding season, the petrels fly from the
Antarctic The Antarctic ( or , American English also or ; commonly ) is a polar region around Earth's South Pole, opposite the Arctic region around the North Pole. The Antarctic comprises the continent of Antarctica, the Kerguelen Plateau and other ...
pack ice Drift ice, also called brash ice, is sea ice that is not attached to the shoreline or any other fixed object (shoals, grounded icebergs, etc.).Leppäranta, M. 2011. The Drift of Sea Ice. Berlin: Springer-Verlag. Unlike fast ice, which is "fasten ...
to the
subtropics The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° north and ...
.


Behaviour


Breeding

Both sexes help to build a nest, and will help incubate the egg. Upon hatching, both sexes again will assist in feeding and protecting the young.Maynard, B. J. (2003)


Feeding

Their diet is composed mainly of
krill Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in n ...
followed by fish. White-chinned petrels feed by surface seizing and by undertaking shallow dives, and they will readily follow ships to collect fisheries discards,Cherel, Y, et al. (1996) making them vulnerable to
longline Long line or longline may refer to: *''Long Line'', an album by Peter Wolf *Long line (topology), or Alexandroff line, a topological space *Long line (telecommunications), a transmission line in a long-distance communications network *Longline fish ...
fisheries.


Conservation

A 2004 estimate placed the adult population at 7,000,000 with an occurrence range of . Mortality of both adults and chicks due to
longline Long line or longline may refer to: *''Long Line'', an album by Peter Wolf *Long line (topology), or Alexandroff line, a topological space *Long line (telecommunications), a transmission line in a long-distance communications network *Longline fish ...
fisheries has caused the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
to classify it as vulnerable. An overall decline in population is inferred by a drop in
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
occupancy rates of 28% at Bird Island, and an 86% reduction in population at
Prydz Bay Prydz Bay is a deep embayment of Antarctica between the Lars Christensen Coast and Ingrid Christensen Coast. The Bay is at the downstream end of a giant glacial drainage system that originates in the East Antarctic interior. The Lambert Glacie ...
. Also, monitoring on
Marion Island The Prince Edward Islands are two small uninhabited islands in the sub-Antarctic Indian Ocean that are part of South Africa. The islands are named Marion Island (named after Marc-Joseph Marion du Fresne, 1724–1772) and Prince Edward Island ...
has shown of 14.5% reduction, and a 37% reduction on the
Crozet Islands The Crozet Islands (french: Îles Crozet; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarcti ...
. Unintentional death at the hands of longline fisheries has proven to be a major contributor to the overall population decline. Nearly all of the
bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
from the
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
n
hake The term hake refers to fish in the: * Family Merlucciidae of northern and southern oceans * Family Phycidae (sometimes considered the subfamily Phycinae in the family Gadidae) of the northern oceans Hake Hake is in the same taxonomic order (Gad ...
are white-chinned petrels. 10% of the
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
n
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
longline bycatch and 55% of the
demersal The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer of ...
. Also, the white-chinned petrel has suffered at the hands of invasive species such as the
brown rat The brown rat (''Rattus norvegicus''), also known as the common rat, street rat, sewer rat, wharf rat, Hanover rat, Norway rat, Norwegian rat and Parisian rat, is a widespread species of common rat. One of the largest muroids, it is a brown o ...
, ''Rattus novegicus'', and the
black rat The black rat (''Rattus rattus''), also known as the roof rat, ship rat, or house rat, is a common long-tailed rodent of the stereotypical rat genus ''Rattus'', in the subfamily Murinae. It likely originated in the Indian subcontinent, but is n ...
, ''Rattus rattus''.BirdLife International (2009) Several of the breeding islands are protected areas. Ongoing studies and population monitoring are occurring at
South Georgia South Georgia ( es, Isla San Pedro) is an island in the South Atlantic Ocean that is part of the British Overseas Territory of South Georgia and the South Sandwich Islands. It lies around east of the Falkland Islands. Stretching in the east ...
, the
Prince Edward Islands The Prince Edward Islands are two small uninhabited islands in the sub-Antarctic Indian Ocean that are part of South Africa. The islands are named Marion Island (named after Marc-Joseph Marion du Fresne, 1724–1772) and Prince Edward Island ...
,
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the sub-Antarctic constituting one of the two exposed parts of the Kerguelen Plateau, a large ...
, and
Crozet Islands The Crozet Islands (french: Îles Crozet; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarcti ...
. Finally, they are a part of ACAP Annex I, and CMS Appendix II. To assist in reversing the decline in population it has been proposed to continue and extend monitoring studies, trying to eliminate most invasive species, promote adoption of best practice mitigation measures at all fisheries within the range via ACAP,
FAO The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
,
CCAMLR The Convention on the Conservation of Antarctic Marine Living Resources, also known as the Commission for the Conservation of Antarctic Marine Living Resources, and CCAMLR, is part of the Antarctic Treaty System. The convention was opened for s ...
.


References


Sources

* * * * * * * *


External links


BirdLife species factsheet
* Whitechinned Petrel
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q276206
white-chinned petrel The white-chinned petrel (''Procellaria aequinoctialis'') also known as the Cape hen and shoemaker, is a large shearwater in the family Procellariidae. It ranges around the Southern Ocean as far north as southern Australia, Peru and Namibia, and ...
white-chinned petrel The white-chinned petrel (''Procellaria aequinoctialis'') also known as the Cape hen and shoemaker, is a large shearwater in the family Procellariidae. It ranges around the Southern Ocean as far north as southern Australia, Peru and Namibia, and ...
Birds of the Southern Ocean Birds of South Australia Birds of New Zealand Birds of Southern Africa Birds of islands of the Atlantic Ocean Birds of subantarctic islands Vulnerable fauna of Australia Fauna of the Crozet Islands Birds of the Indian Ocean Fauna of the Prince Edward Islands
white-chinned petrel The white-chinned petrel (''Procellaria aequinoctialis'') also known as the Cape hen and shoemaker, is a large shearwater in the family Procellariidae. It ranges around the Southern Ocean as far north as southern Australia, Peru and Namibia, and ...
white-chinned petrel The white-chinned petrel (''Procellaria aequinoctialis'') also known as the Cape hen and shoemaker, is a large shearwater in the family Procellariidae. It ranges around the Southern Ocean as far north as southern Australia, Peru and Namibia, and ...