''Procaimanoidea'' ("Before

Caiman A caiman (also cayman as a variant spelling) is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family, the other being alligators. Caimans inhabit Mexico, Central and South America fro ...

-forms") is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

alligatorid The family (biology), family Alligatoridae of crocodylians includes alligators, caimans and their extinct relatives. Phylogeny The superfamily Alligatoroidea includes all crocodilians (fossil and extant) that are more closely related to the Am ...

from the

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

. It was named posthumously in 1946 by

Charles W. Gilmore Charles Whitney Gilmore (March 11, 1874 – September 27, 1945) was an American paleontologist who gained renown in the early 20th century for his work on vertebrate fossils during his career at the United States National Museum (now the N ...

; the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''P. utahensis'', from the

Uintan The Uintan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 46,200,000 to 42,000,000 years BP lasting . It falls within the ...

(middle Eocene) of

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

. It is based on

USNM The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7. ...

15996, a nearly complete skull and partial left hind leg. A second species, ''P. kayi'', was named in 1941 by C.C. Mook as a species of ''

Hassiacosuchus ''Hassiacosuchus'' is an extinct genus of small alligatorid from the early Eocene of Germany, found at the Messel pit. It was named in 1935 by K. Weitzel, and the type species is ''H. haupti''. A second species, ''H. kayi'', was named in 1941 by ...

'', for remains from the

Bridgerian The Bridgerian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 50,300,000 to 46,200,000 years BP lasting . It is usually c ...

(early Eocene) of

Wyoming Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the south ...

. It was reassigned to ''Procaimanoidea'' in 1967 by Wassersug and Hecht.

Description

''Procaimanoidea'' was a small alligatorid, and slightly

heterodont In anatomy, a heterodont (from Greek, meaning 'different teeth') is an animal which possesses more than a single tooth morphology. In vertebrates, heterodont pertains to animals where teeth are differentiated into different forms. For example, ...

, the last four teeth on each side of the jaws having blunt tips.

Phylogeny

Recent studies have consistently resolved ''Procaimanoidea'' as a member of

Alligatorinae Alligatorinae is a subfamily within the family Alligatoridae that contains the alligators and their closest extinct relatives, and is the sister taxon to Caimaninae (the caimans). Many genera in Alligatorinae are described, but only the genus ' ...

, although its relative placement is disputed, as shown by the

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

s below. Cladogram from 2018 Bona ''et al.'' study: Cladogram from 2019 Massonne ''et al.'' study: Cladogram from 2020 Cossette & Brochu study:

References

External links

''Procaimanoidea''
in the

Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) originated in the NCEAS-funded Phanerozoic Marine Pale ...

Alligatoridae Eocene crocodylomorphs Eocene reptiles of North America Prehistoric pseudosuchian genera Taxa named by Charles W. Gilmore {{paleo-archosaur-stub