''Hemipilia lepida'' is a species of

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...

in the family

Orchidaceae Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...

found on

Kyushu Island is the third-largest island of Japan's five main islands and the most southerly of the four largest islands ( i.e. excluding Okinawa). In the past, it has been known as , and . The historical regional name referred to Kyushu and its surroun ...

in Japan, and in the

Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Taiwan: the Ōsumi, Tokara, Amami, Okinawa, and Sakishima Islands (further divided into the Miyako and Yaeyama Islands), with Yonaguni ...

to the south.

Taxonomy

The species was first described by Heinrich Gustav Reichenbach in 1878, as ''Gymnadenia lepida''. It has been placed in various genera, including ''

Gymnadenia ''Gymnadenia'' is a genus of flowering plants in the orchid family (Orchidaceae) containing 22 terrestrial species. The former genus ''Nigritella'' is now included in ''Gymnadenia''. They can be found in damp meadows, fens and marshes, and on ch ...

'', '' Orchis'' and ''Amitostigma''. A

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study in 2014, in which it was included as ''Amitostigma lepidum'', found that species of ''Amitostigma'', ''Neottianthe'' and ''Ponerorchis'' were mixed together in a single

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

, making none of the three genera

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

as then

circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ...

. ''Amitostigma'' and ''Neottianthe'' were subsumed into ''Ponerorchis'', with this species then becoming ''Ponerorchis lepida''. The genus ''Ponerorchis'' has since been synonymized with the genus ''Hemipilia'', resulting in the present name.

References

lepida Orchids of Japan Flora of the Ryukyu Islands Plants described in 1878 {{Orchidoideae-stub