''Ponerorchis keiskei'' (

synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

''Amitostigma keiskei'') is a species of

plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...

in the family

Orchidaceae Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...

. The species is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

Taxonomy

The species was first described in 1879 by Karl Maximovich, using the name "''Gymnadenia keiskei''". However, this name had already been published in 1878 for a different orchid species, so Maximovich's name was illegitimate. In 1900,

Achille Eugène Finet Achille Eugène Finet (1863, Argenteuil – 1913, París) was a French botanist best known for his study of orchids native to Japan and China. Within the family Orchidaceae, he was the taxonomic authority of the genera '' Arethusantha'', ...

reduced the species to a variety as ''Gymnadenia gracilis'' var. ''keiskei'' – this was the first legitimate name. In 1919, Friedrich Rudolf Schlechter transferred the species to ''Amitostigma'' as ''Amitostigma keiskei''. A

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study in 2014 included ''Amitostigma keiskei'' and found that species of ''Amitostigma'', ''Neottianthe'' and ''

Ponerorchis ''Ponerorchis'' is a genus of Asian terrestrial tuberous orchids (family Orchidaceae). It is native to temperate Eurasia, from Poland to Japan, to the north of the Indian subcontinent and to northern Indochina. They can be found in evergreen for ...

'' were mixed together in a single

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

, making none of the three genera

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

as then

circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ...

. ''Amitostigma'' and ''Neottianthe'' were subsumed into ''Ponerorchis'', with this species becoming ''Ponerorchis keiskei''.

References

External links

*
Biglobe, イワチドリ, 岩千鳥 , オオイワチドリ, 大岩千鳥 ''Amitostigma keiskei''Hardy Terrestrial Orchids, Brooklyn Botanical Garden, ''Amitostigma keiskei''
keiskei Orchids of Japan Plants described in 1900 {{Orchidoideae-stub