''Pomacea columellaris'' is a

South American South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

species of

freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...

in the apple snail family, ''

Ampullariidae Ampullariidae, commonly known as the apple snails, is a family of large freshwater snails, aquatic gastropod mollusks with a gill and an operculum. These snails simultaneously have a gill and a lung as functional respiratory structures, wh ...

''.

Taxonomy

''Pomacea columellaris'' was originally described as ''Ampullaria columellaris'' by

Augustus Addison Gould Augustus Addison Gould (April 23, 1805 – September 15, 1866) was an American conchologist and malacologist. Biography Born in New Ipswich, New Hampshire, he was the son of music teacher Nathaniel Duren Gould (1781–1864) who was also noted ...

in 1848, based on a

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

shell collected during the

United States Exploring Expedition The United States Exploring Expedition of 1838–1842 was an exploring and surveying expedition of the Pacific Ocean and surrounding lands conducted by the United States. The original appointed commanding officer was Commodore Thomas ap Catesby ...

from 1838-1842. They were named for their notable

columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved and forms an important source on Roman agriculture, together with the wo ...

, comparable to those of ''

Helix A helix () is a shape like a corkscrew or spiral staircase. It is a type of smooth space curve with tangent lines at a constant angle to a fixed axis. Helices are important in biology, as the DNA molecule is formed as two intertwined helices, ...

'' land snails. In 1904, Dall proposed a

section Section, Sectioning or Sectioned may refer to: Arts, entertainment and media * Section (music), a complete, but not independent, musical idea * Section (typography), a subdivision, especially of a chapter, in books and documents ** Section sign ...

subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...

of '' Ampullaria'' (later treated as

synonymous A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

with ''

Pomacea ''Pomacea'' is a genus of freshwater snails with gills and an operculum, aquatic gastropod mollusks in the family Ampullariidae, the apple snails. The genus is native to the Americas; most species in this genus are restricted to South America. ...

'') called '' Limnopomus'', with ''A. columellaris'' as the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

. After 1991, ''Limnopomus'' was also considered synonymous with ''Pomacea''.

Shell description

The species has a heavy, oval shell with a sharp

spire A spire is a tall, slender, pointed structure on top of a roof of a building or tower, especially at the summit of church steeples. A spire may have a square, circular, or polygonal plan, with a roughly conical or pyramidal shape. Spires are ...

. Its operculum is

corneous Corneous is a biological and medical term meaning horny, in other words made out of a substance similar to that of horns and hooves in some mammals. The word is generally used to describe natural or pathological anatomical structures made out o ...

and able to retract inside the shell's

aperture In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. An opt ...

. They lack an umbilicus and are often yellow in color.

Distribution

''P. columellaris'' is found in rivers in the highlands of

Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...

Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...

(

Pastaza Province Pastaza () is a province in the Oriente of Ecuador located in the eastern jungle. The capital is Puyo, founded on May 12, 1899 and which has 36,700 inhabitants. The city is now accessible by paved roads, a recent development; the main road from ...

), and

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

References

* Simone, L. R. L. (2006). Land and Freshwater Molluscs of Brazil. Editora Grafíca Bernardi, FAPESP. São Paulo, 390 pp

External links

Gould, A.A. (1848). [Shells collected by the United States Exploring Expedition under the command of Charles Wilkes.
Proceedings of the Boston Society of Natural History. 3: 73–75]
Philippi, R. A. (1851-1852). Die Gattung Ampullaria. In Abbildungen nach der Natur mit Beschreibungen. In: Schuberth, G. H.; Wagner, J. A., Eds. Systematisches Conchylien-Cabinet von Martini und Chemnitz. Neu herausgegeben und vervollständigt. Ersten Bandes zwanzigste Abtheilung. 1-74, pls A, 1-21. Nürnberg: Bauer & Raspe
{{Taxonbar, from=Q108528857 columellaris Gastropods described in 1848 Freshwater snails Molluscs of South America Invertebrates of Ecuador Invertebrates of Peru Molluscs of Venezuela Invertebrates of Colombia