''Polypodium scouleri'' is a species of

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

known by the common names leathery polypody, Scouler's polypody, coast polypody and leather-leaf fern. It is native to coastal western North America from

British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...

Guadalupe Island Guadalupe Island ( es, Isla Guadalupe, link=no) is a volcanic island located off the western coast of Mexico's Baja California Peninsula and about southwest of the city of Ensenada in the state of Baja California, in the Pacific Ocean. The ...

off

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

. It is a plant of the coastline, growing in cracks on coastal bluffs, in oceanside forests, beach

dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, f ...

s, and similar habitat. It is often affected by heavy

fog Fog is a visible aerosol consisting of tiny water droplets or ice crystals suspended in the air at or near the Earth's surface. Reprint from Fog can be considered a type of low-lying cloud usually resembling stratus, and is heavily influ ...

s and

sea spray Sea spray are aerosol particles formed from the ocean, mostly by ejection into Earth's atmosphere by bursting bubbles at the air-sea interface. Sea spray contains both organic matter and inorganic salts that form sea salt aerosol (SSA). SSA ha ...

. This

polypody ''Polypodium'' is a genus of ferns in the family Polypodiaceae, subfamily Polypodioideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). The genus is widely distributed throughout the world, with the highest s ...

anchors with a waxy, scaly

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...

. It produces triangular or oblong leaves up to 85 centimeters in maximum length and 8 cm width. Each leaf is made up of many round-tipped linear or oblong segments which are usually stiff and leathery in texture and edged with shallow, rounded teeth. The underside of each leaf segment is crowded with rounded sori each up to half a centimeter wide. The sori contain the

spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, f ...

References

Jepson
* C.Michael Hogan, ed. 2010
‘’Polypodium scouleri’’ . Encyclopedia of Life.

External links

Jepson Manual Treatment — ''Polypodium scouleri''Washington Burke Museum''Polypodium scouleri'' - Photo gallery
scouleri Ferns of the United States Ferns of California Flora of the West Coast of the United States Flora of Baja California Flora of Mexican Pacific Islands Natural history of the California chaparral and woodlands Natural history of the California Coast Ranges Natural history of the San Francisco Bay Area Ferns of the Americas {{Polypodiaceae-stub