''Podarcis carbonelli'', commonly known as Carbonell's wall lizard, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Lacertidae The Lacertidae are the family (biology), family of the wall lizards, true lizards, or sometimes simply lacertas, which are native to Afro-Eurasia. It is a diverse family with at least 300 species in 39 genera. They represent the dominant group o ...

. The species is native to

Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...

and

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

. This lizard reaches a total length (including tail) of 20 cm (8 in), and feeds primarily on small

invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

such as

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

arachnid Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vinegaroon ...

s, and

snails A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastrop ...

. Its natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s are temperate

shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...

s and

sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class of s ...

shore A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past a ...

Habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

threatens its survival.

Etymology

The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''carbonelli'', is in honor of "J. Carbonell" who is the wife of Pérez-Mellado. Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Podarcis carbonelli'', p. 47).

Description

Carbonell's wall lizard grows to a snout-to-vent length of with a tail about twice as long. Females tend to be slightly larger than males in some localities. The dorsal surface is usually grey or brown, but is sometimes green (especially so in males), copiously speckled with rows of dark markings. The flanks may also be somewhat greenish with reticulated, dark markings. The underparts are whitish and there are often small blue spots along the edge of the belly. It differs from all other Iberian Wall Lizards by generally showing greenish flanks but no green backs. Carbonell's wall lizard much resembles the closely related

Bocage's wall lizard Bocage's wall lizard (''Podarcis bocagei'') is a species of lizard in the family Lacertidae. The species is endemic to the Iberian Peninsula. Its natural habitats are temperate forests, temperate shrubland, Mediterranean-type shrubby vegetation, s ...

, but that species tends to have more clearly defined markings and a yellow, orange or pink belly, and shows no green flanks and no blue outer ventral scales.

Distribution and habitat

Carbonell's wall lizard is native to Portugal and Spain. Its geographic range consists of a number of isolated populations in western and central Portugal, another in Coto Doñana in southwestern Spain and a separate subspecies is present on the Berlenga Islands off the coast of Portugal. Being specialized on Mediterranean climates with cool to warm summers, it has stable populations at the Atlantic coast, occupying

sand dunes A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, fl ...

with barely any vegetation. However, the remaining inland populations in the western mountainous

Sistema Central The Central System, Spanish and pt, Sistema Central, is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel an ...

, where it inhabits up to , seem to rapidly decrease, probably a result of global warming.

Behaviour

Carbonell's wall lizard is often seen on dry banks where it may be present in large numbers. It takes refuge in cracks and among tree roots.

Diet

''P. carbonelli'' feeds mainly on

arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

s but, particularly on the Berlenga Islands, also consumes snails.

Reproduction

''P. carbonelli'' is

oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...

. In central Portugal there is usually one

clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...

of two eggs each year, but in the Berlengas, several clutches of up to four eggs are laid. These take ten to fifteen weeks to hatch.

Status

Carbonell's wall lizard has a number of isolated populations and its total geographic range is less than . It lives in oak

woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...

and

and although some populations are in protected areas, others are subject to habitat degradation. Although it is common in some suitable habitats, in general the population is thought to be declining and the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

has assessed it as being an "

endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...