''Plectrohyla hartwegi'' (common name: Hartweg's spikethumb frog) is a species of

frog A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...

in the family

Hylidae Hylidae is a wide-ranging family of frogs commonly referred to as "tree frogs and their allies". However, the hylids include a diversity of frog species, many of which do not live in trees, but are terrestrial or semiaquatic. Taxonomy and s ...

. It is found in the

Sierra Madre de Chiapas The Sierra Madre de Chiapas is a major mountain range in Central America. It crosses El Salvador, Guatemala, Mexico and Honduras. The Sierra Madre de Chiapas is part of the American Cordillera, a chain of mountain ranges that consists of an almos ...

and eastern

Oaxaca Oaxaca ( , also , , from nci, Huāxyacac ), officially the Free and Sovereign State of Oaxaca ( es, Estado Libre y Soberano de Oaxaca), is one of the 32 states that compose the political divisions of Mexico, Federative Entities of Mexico. It is ...

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

Sierra de los Cuchumatanes The Sierra de los Cuchumatanes is the highest non-volcanic mountain range in Central America. Its elevations range from to over , and it covers a total area of .Lovell 2005:11 With an area of situated above , it is also the most extensive hig ...

in western

Guatemala Guatemala ( ; ), officially the Republic of Guatemala ( es, República de Guatemala, links=no), is a country in Central America. It is bordered to the north and west by Mexico; to the northeast by Belize and the Caribbean; to the east by H ...

as well as

Sierra de las Minas Sierra (Spanish for "mountain range" and "saw", from Latin '' serra'') may refer to the following: Places Mountains and mountain ranges * Sierra de Juárez, a mountain range in Baja California, Mexico * Sierra de las Nieves, a mountain range i ...

in eastern Guatemala, and Sierra de Omoa in southwestern

Honduras Honduras, officially the Republic of Honduras, is a country in Central America. The republic of Honduras is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Oce ...

. It might be a composite of two or more species.

Etymology and history

This species is named after Norman Hartweg, American herpetologist. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

, collected by

Eizi Matuda (1894–1978) was a Mexican botanist of Japanese origin. In scholarly works his name is generally romanised Romanization or romanisation, in linguistics, is the conversion of text from a different writing system to the Roman (Latin) scri ...

in 1941, was sent to Hartweg, who recognized it as unique but did not want to

describe Shneur Hasofer is a Hasidic musician known as DeScribe. Hasofer's musical style has been characterized as "Hasidic hip-hop," "Hasidic rap" and "Hasidic R&B". Background Hasofer was born to a Chabad Hasidic family in Melbourne, Australia. Hasof ...

a new species based on just one specimen. The species was eventually described in 1968 by , after further specimens had become available.

Description

The

type series In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...

consisted of three specimens, an adult male (holotype) measuring in snout–vent length (SVL), and two subadult males measuring SVL. Based on further specimens, adult males reach SVL and females SVL. The body is robust. The head is slightly wider than long and as wide as the body; the snout is short. The eyes are large. The heavy supra-tympanic fold covers the upper edge of the tympanum. The

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...

coloration varies is dull olive-brown, olive-green, or pale green. About one fifth of individuals have irregular pale bronze-tan spots on the dorsum and the limbs. The prepollex (the "spikethumb") in males is massive and bifid.

Habitat and conservation

Its natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s are high-elevation (

above sea level Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''. The comb ...

)

cloud forest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud c ...

s near rocky mountain streams, its breeding habitat. It has disappeared from some locations and appears to be on decline. The reasons are probably

chytridiomycosis Chytridiomycosis ( ) is an infectious disease in amphibians, caused by the chytrid fungi ''Batrachochytrium dendrobatidis'' and ''Batrachochytrium salamandrivorans''. Chytridiomycosis has been linked to dramatic population declines or extincti ...

as well as

habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

. The Mexican law protects it under the "Special Protection" category.

References

{{Taxonbar, from=Q917959 hartwegi Frogs of North America Amphibians of Guatemala Amphibians of Honduras Amphibians of Mexico Critically endangered fauna of North America Amphibians described in 1968 Taxa named by William Edward Duellman Taxonomy articles created by Polbot