Pleasing Fungus Beetle
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''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s belonging to
Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark ...
containing over 100
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
. In the present circumscription, it contains 6 tribes (
Tritomini Tritomini is a tribe of pleasing fungus beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present cir ...
, Dacnini, Megalodacnini, Erotylini, Cryptophilini, and
Languriini Languriini is a tribe of lizard beetles in the family Erotylidae, subfamily Languriinae. There are about 5 genera and at least 20 described species in Languriini. ITIS Taxonomic note: *Authorship corrected to Hope 1840, per Bouchard et al., (20 ...
) and 10 subfamilies ( Cryptophilinae, Dacninae, Encaustinae,
Erotylinae Erotylinae is a subfamily of pleasing fungus beetles in the family Erotylidae. Selected genera * ''Acropteroxys'' Gorham, 1887 * '' Antillengis'' * '' Aegithus'' * '' Altisessor'' * '' Aulacochilus'' * ''Bacis'' * '' Brachysphaenus'' * '' Cocc ...
, Languriinae, Loberinae, Megalodacninae, Pharaxonothinae, Tritominae, and
Xenoscelinae Xenoscelinae is a subfamily of pleasing fungus beetles in the family Erotylidae. There are about 8 genera and 12 described species in Xenoscelinae. Genera These eight genera belong to the subfamily Xenoscelinae: * '' Cryptophilus'' Reitter, 187 ...
). In other words, the narrowly circumscribed Erotylidae correspond to the subfamily Erotylinae in the definition ''
sensu lato ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
''. There are doubts on the
monophyly In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic grou ...
of lower ranked taxa within Erotylidae, with further phylogenetic studies requiring better sampling and studies of unexplored character sets, for example the metendosternite and penile flagellum, which are generally lacking detailed morphological studies within the Coleoptera literature. The Eroytlina taxonomy is based off traits such as their different colors and not off morphological differences like mouthparts, thorax, and abdominal terminalia (Pecci-Maddalena). Erotylidae feed on
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
and
fungal A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from th ...
matter; some are important
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
s (e.g. of the ancient
cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...
s), while a few have gained notoriety as pests of some significance. Sometimes, useful and harmful species are found in one
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
, e.g. ''
Pharaxonotha ''Pharaxonotha'' is a genus of pleasing fungus beetles in the family Erotylidae. There are at least three described species in ''Pharaxonotha''. Species These three species belong to the genus ''Pharaxonotha'': * '' Pharaxonotha cerradensis'' S ...
''. Most pleasing fungus beetles, however, are inoffensive animals of little significance to humans. The oldests fossil is an undescribed specimen known from Early Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
)
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and southwest Syria. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropi ...
.


Selected genera

These 160 genera belong to the family Erotylidae: * ''
Acropteroxys ''Acropteroxys'' is a genus of lizard beetles in the family Erotylidae. There are at least two described species in ''Acropteroxys''. Species These two species belong to the genus ''Acropteroxys'': * ''Acropteroxys gracilis'' (Newman, 1838) (sl ...
'' Gorham, 1887 i c g b * '' Acryptophagus'' * '' Aegithus'' Fabricius, 1801 g * '' Amblyopus'' Lacordaire, 1842 g * '' Amblyscelis'' Gorham, 1888 * '' Anadastus'' Gorham, 1887 g * '' Apolybas'' Alvarenga, 1965 g * '' Atomarops'' Reitter * '' Aulacochilus'' Lacordaire, 1842 * '' Bacis'' Dejean, 1836 g * '' Barytopus'' Chevrolat, 1836 g * '' Bolerus'' Grouvelle * '' Brachypterosa'' Zablotny & Leschen, 1996 * '' Brachysphaenus'' * '' Caenolanguria'' Gorham, 1887 g * '' Callischyrus'' * '' Camptocarpus'' * '' Cathartocryptus'' Sharp, 1886 i c g * '' Chinophagus'' Ljubarsky, 1997 * '' Cladoxena'' Motschulsky * '' Cnecosa'' * '' Coccimorphus'' Hope, 1841 * '' Coelocryptus'' Sharp, 1900 * '' Combocerus'' Bedel, 1868 g * '' Coptengis'' Crotch, 1876 * '' Crotchia'' Fowler * '' Crowsenguptus'' * '' Cryptodacne'' Sharp, 1878 g * ''
Cryptophilus ''Cryptophilus'' is a genus of pleasing fungus beetles in the family Erotylidae. There are about nine described species in ''Cryptophilus''. Identification ''Cryptophilus'' adults are brown and slightly hairy beetles, 1.6-2.3 mm long and with ...
'' Reitter, 1874 i c g b * '' Cycadophila'' Xu, Tang & Skelley, 2015 g * '' Cypherotylus'' Crotch, 1873 * '' Cyrtomorphus'' Lacordaire, 1842 g * '' Cytorea'' * '' Dacne'' Latrielle, 1796 * '' Dactylotritoma'' Arrow, 1925 g * '' Dapsa'' * ''
Dasydactylus ''Dasydactylus'' is a genus of lizard beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circum ...
'' Gorham, 1887 i c g b * '' Doubledaya'' White, 1850 g * '' Ectrapezidera'' * '' Ellipticus'' Chevrolat, 1836 g * '' Empocryptus'' Sharp * '' Encaustes'' Lacordaire, 1842 g * '' Epilanguria'' Fowler, 1908 g * ''
Episcapha ''Episcapha'' is a genus of beetles belonging to the family Salpingidae Salpingidae or narrow-waisted bark beetles is a family of beetles in the superfamily Tenebrionoidea. The species are small, about 1.5 – 7 mm in length. The famil ...
'' Dejean, 1837 * '' Episcaphula'' Crotch * '' Erotylina'' Curran, 1944 g * '' Erotylus'' Fabricius, 1775 * '' Eutriplax'' Lewis, 1887 * '' Fitoa'' * '' Goniolanguria'' * '' Haematochiton'' Gorham, 1888 i c g b * ''
Hapalips ''Hapalips'' is a genus of pleasing fungus beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present c ...
'' Reitter, 1877 i c g b * '' Henoticonus'' Reitter, 1878 * '' Hirsotriplax'' * '' Hirsutotriplax'' Skelley, 1993 i c g b * '' Homoeotelus'' Hope, 1841 * '' Hoplepiscapha'' Lea, 1922 * '' Hornerotylus'' g * ''
Iphiclus ''Iphiclus'' is a genus of beetles in the family Erotylidae. It was first described by Louis Alexandre Auguste Chevrolat Louis Alexandre Auguste Chevrolat was a French entomologist, born 29 March 1799 in Paris and died 16 December 1884 in Pa ...
'' Chevrolat, 1836 g * '' Ischyrus'' Lacordaire, 1842 i c g b * ''
Languria ''Languria'' is a genus of lizard beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circumscri ...
'' Latreille, 1802 i c g b * '' Languriomorpha'' * ''
Langurites ''Langurites'' is a genus of lizard beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circumsc ...
'' Motschulsky, 1860 i c g b * '' Lepidotoramus'' Leschen, 1997 i c g * '' Leucohimatium'' Rosenhauer, 1856 * '' Ligurana'' Chûjô, 1974 g * '' Linodesmus'' Bedel, 1882 * '' Loberogosmus'' Reitter * '' Loberolus'' * '' Loberonotha'' Sen Gupta and Crowson, 1969 i c g * '' Loberopsyllus'' * '' Loberoschema'' Reitter * ''
Loberus ''Loberus'' is a genus of pleasing fungus beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present ci ...
'' LeConte, 1861 i c g b * '' Lobosternum'' Reitter, 1875 i c g * '' Lybanodes'' * '' Lybas'' Lacordaire, 1842 g * '' Macromelea'' Hope * '' Macrophagus'' Motschulsky * '' Malleolanguria'' * '' Megalodacne'' Crotch, 1873 i c g b * '' Megischyrus'' Crotch, 1873 g * '' Meristobelus'' * '' Micrencaustes'' Crotch, 1875 g * '' Microlanguria'' Lewis * '' Microsternus'' Lewis, 1887 i c g b * '' Mimodacne'' * '' Mycetaea'' Stephens, 1829 * '' Mycolybas'' Crotch, 1876 * '' Mycophtorus'' Lacordaire, 1842 g * ''
Mycotretus ''Mycotretus'' is a genus of pleasing fungus beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present ...
'' Lacordaire, 1842 i c g b * '' Neocladoxena'' Maeda, 1974 g * '' Neodacne'' Chûjô, 1976 g * '' Neoloberolus'' Leschen, 2003 i c g * '' Neopriotelus'' Alvarenga, 1965 g * '' Neosternus'' Dai & Zhao, 2013 g * '' Neotriplax'' * '' Neoxestus'' Crotch, 1875 g * '' Nomotus'' * '' Oligocorynus'' Chevrolat, 1836 g * '' Oretylus'' Heller, 1920 g * '' Ortholanguria'' * '' Othniocryptus'' Sharp, 1900 * '' Pachylanguria'' Crotch, 1875 g * '' Paederolanguria'' Mader, 1939 g * '' Paphezia'' Zablotny & Leschen, 1996 * '' Paracladoxena'' Fowler * '' Paraxonotha'' g * '' Pediacus'' * '' Penolanguria'' Kolbe * ''
Pharaxonotha ''Pharaxonotha'' is a genus of pleasing fungus beetles in the family Erotylidae. There are at least three described species in ''Pharaxonotha''. Species These three species belong to the genus ''Pharaxonotha'': * '' Pharaxonotha cerradensis'' S ...
'' Reitter, 1875 i c g b * '' Phricobacis'' Crotch, 1876 g * '' Platoberus'' Sharp * '' Prepopharus'' Erichson, 1847 g * '' Promecolanguria'' * '' Protoloberus'' Leschen, 2003 i c g * '' Pselaphacus'' Percheron, 1835 g * '' Pselaphandra'' Jacobson, 1904 * '' Pseudhapalips'' Champion * '' Pseudhenoticus'' Sharp * '' Pseudischyrus'' Casey, 1916 i c g b * '' Pseudotritoma'' Gorham, 1888 g * '' Rhodotritoma'' Arrow, 1925 g * '' Scaphengis'' Gorham, 1888 g * '' Scaphidomorphus'' Hope, 1841 g * '' Scaphodacne'' Heller, 1918 * '' Scelidopetalon'' Delkeskamp, 1957 g * '' Setariola'' Jakobson, 1915 g * '' Sphenoxus'' Lacordaire, 1842 g * '' Spondotriplax'' Crotch, 1875 g * '' Stengita'' * '' Stenodina'' * '' Strongylosomus'' Chevrolat, 1836 g * '' Tapinotarsus'' Kirsch, 1865 g * '' Telmatoscius''* '' Teretilanguria'' * '' Tetralanguria'' Crotch, 1875 g * '' Tetraphala'' g * '' Tetratritoma'' Arrow, 1925 g * '' Thallis'' * '' Thallisella'' Crotch * '' Thallisellodes'' Arrow, 1925 g * '' Tomarops'' Grouvelle, 1903 * ''
Toramus ''Toramus'' is a genus of pleasing fungus beetles in the family Erotylidae. There are about nine described species in ''Toramus''. Species These nine species belong to the genus ''Toramus'': * '' Toramus acutus'' (Reitter, 1875) * '' Toramus bis ...
'' Grouvelle, 1916 i c g b * '' Trapezidera'' * '' Trapezidistes'' Fowler, 1887 g * '' Trichotritoma'' * '' Trichulus'' * '' Triplacidea'' Gorham, 1901 * ''
Triplax ''Triplax'' is a genus of pleasing fungus beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present ci ...
'' Herbst, 1793 i c g b * ''
Tritoma ''Tritoma'' is a genus of beetles in the family Erotylidae, the pleasing fungus beetles. It is distributed worldwide, mainly in the Old World.Goodrich, M. A. and C. A. Springer. (1999)The pleasing fungus beetles (Coleoptera: Erotylidae) of Nebr ...
'' Fabricius, 1775 i c g b * '' Truquiella'' b * '' Typocephalus'' Chevrolat, 1837 g * '' Xenocryptus'' Arrow, 1929 * '' Xenohimatium'' Lyubarsky & Perkovsky, 2012 g * '' Xenoscelinus'' Grouvelle, 1910 g * '' Xenoscelis'' Wollaston, 1864 g * '' Xestus'' Wollaston, 1864 g * '' Zavaljus'' Reitter, 1880 g* ''
Acropteroxys ''Acropteroxys'' is a genus of lizard beetles in the family Erotylidae. There are at least two described species in ''Acropteroxys''. Species These two species belong to the genus ''Acropteroxys'': * ''Acropteroxys gracilis'' (Newman, 1838) (sl ...
'' Gorham, 1887 * '' Zonarius'' (= ''Oligocorynus'') * '' Zythonia'' Westwood, 1874 Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net


References

* * * *


External links

* *
Key to the British species of family Erotylidae
{{Taxonbar, from=Q1194119 Cucujoidea families