A planula is the free-swimming, flattened,

ciliated The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projecti ...

, bilaterally symmetric larval form of various

cnidaria Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that th ...

n species and also in some species of

Ctenophores Ctenophora (; ctenophore ; ) comprise a phylum of marine invertebrates, commonly known as comb jellies, that inhabit sea waters worldwide. They are notable for the groups of cilia they use for swimming (commonly referred to as "combs"), a ...

. Some groups of

Nemertea Nemertea is a phylum of animals also known as ribbon worms or proboscis worms, consisting of 1300 known species. Most ribbon worms are very slim, usually only a few millimeters wide, although a few have relatively short but wide bodies. Many h ...

ns also produce larvae that are very similar to the planula, which are called planuliform larva.

Development

The planula forms either from the fertilized egg of a

medusa In Greek mythology, Medusa (; Ancient Greek: Μέδουσα "guardian, protectress"), also called Gorgo, was one of the three monstrous Gorgons, generally described as winged human females with living venomous snakes in place of hair. Those ...

, as is the case in scyphozoans and some

hydrozoa Hydrozoa (hydrozoans; ) are a taxonomic class of individually very small, predatory animals, some solitary and some colonial, most of which inhabit saline water. The colonies of the colonial species can be large, and in some cases the specializ ...

ns, or from a polyp, as in the case of

anthozoa Anthozoa is a subphylum of marine invertebrates which includes the sea anemones, Scleractinia, stony corals and Alcyonacea, soft corals. Adult anthozoans are almost all attached to the seabed, while their larvae can disperse as part of the plank ...

ns. Depending on the species, the planula either metamorphoses directly into a free-swimming, miniature version of the mobile adult form, or navigates through the water until it reaches a hard substrate (many may prefer specific substrates) where it anchors and grows into a polyp. The miniature-adult types include many open-ocean scyphozoans. The attaching types include all

ns with a planula stage, many coastal scyphozoans, and some

ns.

Feeding and locomotion

The planulae of the subphylum

Medusozoa Medusozoa is a clade in the phylum Cnidaria, and is often considered a subphylum. It includes the classes Hydrozoa, Scyphozoa, Staurozoa and Cubozoa, and possibly the parasitic Polypodiozoa. Medusozoans are distinguished by having a medusa sta ...

have no mouth, and no digestive tract, and are unable to feed themselves, while those of

Anthozoa Anthozoa is a subphylum of marine invertebrates which includes the sea anemones, Scleractinia, stony corals and Alcyonacea, soft corals. Adult anthozoans are almost all attached to the seabed, while their larvae can disperse as part of the plank ...

can feed. Planula larvae swim with the

aboral Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

end (the end opposite the mouth) in front.

References

Cnidarian biology Larvae {{cnidarian-stub