''Plantago indica'', commonly known as branched plantain, sand plantain, or black psyllium, is a

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

in the plantain family

Plantaginaceae Plantaginaceae, the plantain family, is a large, diverse family of flowering plants in the order Lamiales that includes common flowers such as snapdragon and foxglove. It is unrelated to the banana-like fruit also called "plantain." In older cl ...

, and is one of a few species in the ''

Plantago ''Plantago'' is a genus of about 200 species of flowering plants in the family Plantaginaceae, commonly called plantains or fleaworts. The common name plantain is shared with the unrelated cooking plantain. Most are herbaceous plants, though a ...

'' genus under the common name

psyllium Psyllium , or ispaghula , is the common name used for several members of the plant genus ''Plantago'' whose seeds are used commercially for the production of mucilage. Psyllium is mainly used as a dietary fiber to relieve symptoms of both constip ...

. The plant is native to parts of

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

, and

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...

, and has been naturalized in many other areas such as

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

and

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

. The plant can be found mostly in dry inland areas, such as those that are sandy, and has also

naturalized Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the in ...

on roadsides and in meadows. The plant is not used broadly as a food source, but has been cultivated for its seeds which serve a medicinal use as a

laxative Laxatives, purgatives, or aperients are substances that loosen stools and increase bowel movements. They are used to treat and prevent constipation. Laxatives vary as to how they work and the side effects they may have. Certain stimulant, lubri ...

Description

''Plantago indica'' is an annual herb with a

taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproo ...

that has an erect stem with leaves that are usually opposite but sometimes in whorls of 3, and elongated internodes between leaf sets. Glandular pubescence is found on the stems, leaves, sepals, and

inflorescences An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...

. The leaves are simple and have a base that is decurrent onto the petiole, an entire or slightly dentate edge, are linear or lanceolate in shape, and can reach 2.5-5cm long and 1-3mm wide. Some leaves are modified into

bracts In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...

between 0.5cm and 2cm which have a cuspidate apex and an orbicular-ovate base. The plant has a densely flowered inflorescence with flowers that have a glabrous, brownish corolla and a calyx with broadly veined, elliptic and obovate sepals. The stamens exsert above the corolla, and the anthers are yellow, ellipsoid, and 1.8-2.2mm in size. The style of the stigma is also well exserted, and the floral parts are hypogynous but extend superior to the ovary. The seeds are contained in capsules of 2. The seeds are 2.5-2.8 mm in size, black or blackish-brown, shiny, ellipsoid, and have a distinct central groove on the inner face.

Ecology

''Plantago indica'' is described as being native to

North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...

Southwest China Southwest China () is a region in the south of the People's Republic of China. Geography Southwest China is a rugged and mountainous region, transitioning between the Tibetan Plateau to the west and the Chinese coastal hills (东南丘陵) and ...

Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia to the north and west, China to the east, Kyrgyzstan to the southeast, Uzbeki ...

Kyrgyzstan Kyrgyzstan,, pronounced or the Kyrgyz Republic, is a landlocked country in Central Asia. Kyrgyzstan is bordered by Kazakhstan to the north, Uzbekistan to the west, Tajikistan to the south, and the People's Republic of China to the east. ...

, and

Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...

. The plant has become broadly naturalized, identified in areas of

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

, and

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...

. The plant is commonly found in sandy areas such as in arid deserts and on sandy beaches, and has also been seen distributed on roadsides, and by railroad tracks.

Uses

The seeds, known as French psyllium or black psyllium, are medicinally cultivated, along with other species under the

common name, such as ''P. ovata'', for use as a laxative for constipation and are also used to treat irritable bowel syndrome (IBS) and diarrhea by extracting the

mucilage Mucilage is a thick, gluey substance produced by nearly all plants and some microorganisms. These microorganisms include protists which use it for their locomotion. The direction of their movement is always opposite to that of the secretion of m ...

from the seed coat. The mucilage from the seeds has also been reported to lower the risk of coronary heart disease.

References

{{Taxonbar, from1=Q25399593, from2=Q165095 indica Biologically-based therapies Medicinal plants Flora of Africa Medicinal plants of Africa Flora of Asia Medicinal plants of Asia Flora of Europe Medicinal plants of Europe Flora of Australia Medicinal plants of Australia Flora of North America Medicinal plants of North America