Hardiness of
plant
Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclu ...
s describes their ability to survive adverse growing conditions. It is usually limited to discussions of
climatic
Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorological ...
adversity. Thus a plant's ability to tolerate
cold
Cold is the presence of low temperature, especially in the atmosphere. In common usage, cold is often a subjective perception. A lower bound to temperature is absolute zero, defined as 0.00K on the Kelvin scale, an absolute thermodynamic ...
,
heat
In thermodynamics, heat is defined as the form of energy crossing the boundary of a thermodynamic system by virtue of a temperature difference across the boundary. A thermodynamic system does not ''contain'' heat. Nevertheless, the term is ...
,
drought
A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D. Jiang, A. Khan, W. Pokam Mba, D. Rosenfeld, J. Tierney, an ...
,
flood
A flood is an overflow of water ( or rarely other fluids) that submerges land that is usually dry. In the sense of "flowing water", the word may also be applied to the inflow of the tide. Floods are an area of study of the discipline hydrol ...
ing, or
wind
Wind is the natural movement of air or other gases relative to a planet's surface. Winds occur on a range of scales, from thunderstorm flows lasting tens of minutes, to local breezes generated by heating of land surfaces and lasting a few ho ...
are typically considered measurements of hardiness. Hardiness of plants is defined by their native extent's geographic location:
longitude
Longitude (, ) is a geographic coordinate that specifies the east– west position of a point on the surface of the Earth, or another celestial body. It is an angular measurement, usually expressed in degrees and denoted by the Greek lette ...
,
latitude
In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north pol ...
and
elevation
The elevation of a geographic location is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational surface (see Geodetic datum § Ver ...
. These attributes are often simplified to a
hardiness zone. In
temperate
In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
latitudes, the term most often describes resistance to cold, or "cold-hardiness", and is generally measured by the lowest temperature a plant can withstand.
Hardiness of a plant is usually divided into two categories: tender, and hardy. (Some sources also use the erroneous terms "Half-hardy" or "Fully hardy".) Tender plants are those killed by freezing temperatures, while hardy plants survive freezing—at least down to certain temperatures, depending on the plant. "Half-hardy" is a term used sometimes in horticulture to describe
bedding
Bedding, also known as bedclothes or bed linen, is the materials laid above the mattress of a bed for hygiene, warmth, protection of the mattress, and decorative effect. Bedding is the removable and washable portion of a human sleeping environm ...
plants which are sown in heat in winter or early spring, and planted outside after all danger of frost has passed. "Fully hardy" usually refers to plants being classified under the Royal Horticultural Society classifications, and can often cause confusion to those not using this method.
Plants vary greatly in their tolerance of growing conditions, and are capable of adaptation to changes in climate on their own to some extent. The
selective breeding
Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant m ...
of
varieties
Variety may refer to:
Arts and entertainment Entertainment formats
* Variety (radio)
* Variety show, in theater and television
Films
* ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont
* ''Variety'' (1935 film), ...
capable of withstanding particular climates forms an important part of
agriculture
Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people t ...
and
horticulture
Horticulture is the branch of agriculture that deals with the art, science, technology, and business of plant cultivation. It includes the cultivation of fruits, vegetables, nuts, seeds, herbs, sprouts, mushrooms, algae, flowers, seaweeds and no ...
. Part of the work of nursery growers of plants consists of
cold hardening
Cold hardening is the physiological and biochemical process by which an organism prepares for cold weather.
Plants
Plants in temperate and polar regions adapt to winter and sub zero temperatures by relocating nutrients from leaves and shoots to ...
, or hardening off their plants, to prepare them for likely conditions in later life.
Winter hardiness
Winter-hardy plants grow during the
winter
Winter is the coldest season of the year in polar and temperate climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Different cultur ...
, or at least remain healthy and
dormant. Apart from hardy
evergreen
In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
s, these include many cultivated plants, including varieties of
cabbage and
broccoli
Broccoli (''Brassica oleracea'' var. ''italica'') is an edible green plant in the cabbage family (family Brassicaceae, genus ''Brassica'') whose large flowering head, stalk and small associated leaves are eaten as a vegetable. Broccoli is cla ...
, and all kinds of
carrot. Some bulbs – such as
tulip
Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...
s – need cold winters to bloom, while others – such as
freesia
''Freesia'' is a genus of herbaceous perennial flowering plants in the family Iridaceae, first described as a genus in 1866 by Christian Friedrich Ecklon (1886) and named after the German botanist and medical practitioner, Friedrich Freese (1795 ...
– can survive a freezing winter. Many domestic plants are assigned a
hardiness zone that specifies the climates in which they can survive.
Winter garden
A winter garden is a kind of garden maintained in wintertime.
History
The origin of the winter garden dates back to the 17th to 19th centuries where European nobility would construct large conservatories that would house tropical and subtro ...
s are dependent upon the cultivation of winter-hardy plants.
Mechanisms
Woody plants survive freezing temperatures by suppressing the formation of ice in living cells or by allowing water to freeze in plant parts that are not affected by ice formation. The common mechanism for woody plants to survive down to –40 °C (–40 °F) is
supercooling
Supercooling, also known as undercooling, is the process of lowering the temperature of a liquid or a gas below its melting point without it becoming a solid. It achieves this in the absence of a seed crystal or nucleus around which a crystal ...
. Woody plants that survive lower temperatures are dehydrating their cells, allowing water to freeze between cell walls and the cells to survive. Plants normally considered hardy may not survive freezing if they are not readily acclimated, which renders them unable to use these mechanisms.
Hardiness ratings
Various hardiness ratings are published. In the
United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
(US), the most widely used is the
U.S. Department of Agriculture
The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of comme ...
(USDA) system of hardiness zones based on average minimum yearly temperatures. This system was developed specifically for the extremely diverse range of conditions in the US, from baking desert to frozen tundra. Another commonly used system is the Sunset Climate Zone system. This system is much more specific to climates (i.e. precipitation, temperature, and humidity based) and less dependent on the yearly minimum.
In contrast the
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the European mainland, continental mainland. It comprises England, Scotlan ...
(UK) and
Western Europe
Western Europe is the western region of Europe. The region's countries and territories vary depending on context.
The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the ancient Mediterranean ...
have an oceanic climate, and experience a narrower range of temperatures that is tempered by the presence of the
Gulf Stream. This results in areas like western Scotland experiencing relatively mild winter conditions and cool summers that enable the growing of some subtropical plants, despite being well to the north of subtropical climate areas. The
Royal Horticultural Society
The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity.
The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
has published a set of hardiness ratings applicable to the UK. The ratings range from H1a to H7. H1a, higher than , applies to tropical plants permanently under glass in heat; while H7, below , applies to very cold-tolerant plants such as heathers. Most outdoor plants in the UK fall within the range H4, (hardy in the average winter) to H5, (hardy in a cold winter). Also, the average minimum temperature in the UK is much warmer than the average minimums in much of the northern US, while the average maximums in the northern US are often much warmer in summer than the UK.
In addition to cold tolerance, plant hardiness has been observed to be linked to how much stress specific plants are undergoing into the winter, or even how fast the onset of cold weather is in a specific year. This means that often stressed plants will exhibit less cold tolerance than plants that have been well maintained. Plants may also die if the winter changes from balmy to exceptionally cold in a short period of time.
Other hardiness
Tolerance to other climatic factors are often omitted from the "hardiness" discussion. This section provides a very brief overview of the topics.
* Heat tolerance may be displayed alongside cold-hardiness, such as in the American Horticultural Society's "Heat Zones". See and
Breeding for heat stress tolerance
Plant breeding is process of development of new cultivars. Plant breeding involves development of varieties for different environmental conditions – some of them are not favorable. Among them, heat stress is one of such factor that reduces the pr ...
.
* For drought tolerance, see and
Breeding for drought stress tolerance
Breeding for drought resistance is the process of breeding plants with the goal of reducing the impact of dehydration on plant growth.
Dehydration stress Crop plants
In nature or crop fields, water is often the most limiting factor for plant gr ...
.
* For flood tolerance, see cited review.
* The mechanisms of wind tolerance are still poorly understood. Wind not only causes mechanical stress, but also makes the plant colder and dryer. Plants may evolutionally adapt to wind by becoming smaller (to avoid wind) or bigger (to resist the wind). Physiologically, the mechanical force also causes a cascade of signals in the living plant.
See also
*
Microclimate
A microclimate (or micro-climate) is a local set of atmospheric conditions that differ from those in the surrounding areas, often with a slight difference but sometimes with a substantial one. The term may refer to areas as small as a few squ ...
References
Bibliography
*
*
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External links
Interactive Version of the 1990 USDA Hardiness Zone Map
Plants by adaptation
Agricultural terminology
Crops
Garden plants
Horticulture
Plant ecology
Trees by climate
Climate zones