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''Pinus rigida'', the pitch pine, is a small-to-medium-sized
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
. It is native to eastern
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, primarily from central Maine south to Georgia and as far west as Kentucky. It is found in environments which other species would find unsuitable for growth, such as acidic, sandy, and low-nutrient soils.


Description

The pitch pine is irregular in shape, but grows to ). Branches are usually twisted, and it does a poor job at self-pruning. The needles are in fascicles (bundles) of three, about in length, and are stout (over broad) and often slightly twisted. The cones are long and oval, with prickles on the scales. Trunks are usually straight with a slight curve, covered in large, thick, irregular plates of bark. Pitch pine has an exceptionally high regenerative ability; if the main trunk is cut or damaged by fire, it can re-sprout using
epicormic shoot An epicormic shoot is a shoot growing from an epicormic bud, which lies underneath the bark of a trunk, stem, or branch of a plant. Epicormic buds lie dormant beneath the bark, their growth suppressed by hormones from active shoots higher up ...
s. This is one of its many adaptations to fire, which also include a thick bark to protect the sensitive cambium layer from heat. Burnt pitch pines often form stunted, twisted trees with multiple trunks as a result of the resprouting. This characteristic makes it a popular species for
bonsai Bonsai ( ja, 盆栽, , tray planting, ) is the Japanese art of growing and training miniature trees in pots, developed from the traditional Chinese art form of ''penjing''. Unlike ''penjing'', which utilizes traditional techniques to produce ...
. Pitch pine is rapid-growing when young, gaining around one foot of height per year under optimal conditions, until growth slows at 50–60 years. By 90 years of age, the amount of annual height gain is minimal. Open-growth trees begin bearing cones in as little as three years, with shade-inhabiting pines taking a few years longer. Cones take two years to mature. Seed dispersal occurs over the fall and winter, and trees cannot self-pollinate. The lifespan of a pitch pine is about 200 years or longer.


Taxonomy

It was given its scientific name, ''Pinus rigida'', by British botanist
Philip Miller Philip Miller FRS (1691 – 18 December 1771) was an English botanist and gardener of Scottish descent. Miller was chief gardener at the Chelsea Physic Garden for nearly 50 years from 1722, and wrote the highly popular ''The Gardeners Dictio ...
. It belongs to the family Pinaceae and the subgenus ''
Pinus A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
'' (formerly ''Diploxylon''), along with other hard pines.


Distribution and habitat

Pitch pine is found mainly in the southern areas of the
northeastern United States The Northeastern United States, also referred to as the Northeast, the East Coast, or the American Northeast, is a geographic region of the United States. It is located on the Atlantic coast of North America, with Canada to its north, the Southe ...
, from coastal
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and north ...
and
Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...
to
Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...
and northern
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
. A few stands occur in southern
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...
and
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
, mostly in two pockets along the St. Lawrence River. It is known as a pioneer species and is often the first tree to vegetate a site after it has been cleared away. It is a climax vegetation type in extreme conditions, but in most cases it is replaced by oaks and other hardwoods. This pine occupies a variety of habitats, from dry, acidic sandy uplands to swampy lowlands, and can survive in very poor conditions. It is the primary tree of the
New Jersey Pine Barrens The New Jersey Pine Barrens, also known as the Pinelands or simply the Pines, is the largest remaining example of the Atlantic coastal pine barrens ecosystem, stretching across more than seven counties of New Jersey. Two other large, contiguou ...
.


Ecology

Pitch pines provide habitat and food for many wildlife species. They are used for cover and nesting by birds such as the
pine warbler The pine warbler (''Setophaga pinus'') is a small songbird of the New World warbler family. Description These birds have white bellies, two white wing bars, dark legs and thin, relatively long pointed bills; they have yellowish 'spectacles' a ...
,
wild turkey The wild turkey (''Meleagris gallopavo'') is an Upland game bird, upland ground bird native to North America, one of two extant species of Turkey (bird), turkey and the heaviest member of the order Galliformes. It is the ancestor to the domestic ...
,
red-cockaded woodpecker The red-cockaded woodpecker (''Leuconotopicus borealis'') is a woodpecker endemic to the southeastern United States. Description The red-cockaded woodpecker is small to mid-sized species, being intermediate in size between North America's two ...
,
great-crested flycatcher The great crested flycatcher (''Myiarchus crinitus'') is a large insect-eating bird of the tyrant flycatcher family. It is the most widespread member of the genus ''Myiarchus'' in North America, and is found over most of the eastern and mid-west ...
,
blue jay The blue jay (''Cyanocitta cristata'') is a passerine bird in the family Corvidae, native to eastern North America. It lives in most of the eastern and central United States; some eastern populations may be migratory. Resident populations are a ...
,
black-capped chickadee The black-capped chickadee (''Poecile atricapillus'') is a small, non-migratory, North American songbird that lives in deciduous and mixed forests. It is a passerine bird in the tit family, the Paridae. It is the state bird of Massachusetts and ...
,
black-and-white warbler The black-and-white warbler (''Mniotilta varia'') is a species of New World warbler, and the only member of its genus, ''Mniotilta''. It breeds in northern and eastern North America and winters in Florida, Central America, and the West Indies ...
,
Nashville warbler The Nashville warbler (''Leiothlypis ruficapilla'') is a small songbird in the New World warbler family, found in North and Central America. It breeds in parts of the northern and western United States and southern Canada, and migrates to winter ...
, and
chestnut-sided warbler The chestnut-sided warbler (''Setophaga pensylvanica'') is a New World warbler. They breed in eastern North America and in southern Canada westwards to the Canadian Prairies. They also breed in the Great Lakes region and in the eastern United St ...
. Deer consume seedlings and new sprouts, and small mammals and birds eat the seeds. This species occasionally hybridizes with other pine species, such as loblolly pine (''
Pinus taeda ''Pinus taeda'', commonly known as loblolly pine, is one of several pines native to the Southeastern United States, from East Texas to Florida, and north to southern New Jersey. The wood industry classifies the species as a southern yellow pine. ...
''), shortleaf pine (''
Pinus echinata ''Pinus echinata'', the shortleaf pine, is a species of pine native to the southeastern United States. Description The tree is variable in form, sometimes straight, sometimes crooked, with an irregular crown. The tree reaches heights of with a ...
''), and pond pine (''
Pinus serotina ''Pinus serotina'', the pond pine, marsh pine or pocosin pine, is a pine tree found along the Southeastern portion of the Atlantic coastal plain of the United States, from southern New Jersey south to Florida and west to southern Alabama. This pi ...
''); the last is treated as a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of pitch pine by some
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
s.


Uses

Pitch pine is not a major timber tree due to the frequency of multiple or crooked trunks, nor is it as fast-growing as other eastern American pines. However, it grows well on unfavorable sites. In the past, it was a major source of pitch and timber for ship building, mine timbers, and railroad ties because the wood's high resin content preserves it from decay. As such, it has also been used for elaborate wood constructions, e. g. radio towers. Pitch pine is currently used mainly for rough construction, pulp, crating, and fuel. However, due to its uneven growth, quantities of high quality can be difficult to obtain, and large lengths of pitch pine can be very costly. Archaeology indicates that the
Iroquois The Iroquois ( or ), officially the Haudenosaunee ( meaning "people of the longhouse"), are an Iroquoian-speaking confederacy of First Nations peoples in northeast North America/ Turtle Island. They were known during the colonial years to ...
, Shinnecock, and
Cherokee The Cherokee (; chr, ᎠᏂᏴᏫᏯᎢ, translit=Aniyvwiyaʔi or Anigiduwagi, or chr, ᏣᎳᎩ, links=no, translit=Tsalagi) are one of the indigenous peoples of the Southeastern Woodlands of the United States. Prior to the 18th century, t ...
all utilized pitch pine. The Iroquois used the pitch to treat rheumatism, burns, cuts, and boils. Pitch also worked as a laxative. A pitch pine poultice was used by both the Iroquois and the Shinnecock to open boils and to treat abscesses. The Cherokee used pitch pine wood in canoe construction and for decorative carvings. Pitch pine is known to cross with pond loblolly and shortleaf pines. One of those crosses is the pitlolly pine (pinus x rigitaeda), a natural hybrid between the loblolly pine and the pitch pine. This hybrid combines the tall size of the loblolly pine and the cold- hardiness of the pitch pine. This hybrid was used as substitute of loblolly pine and has been extensively planted in South Korea.


Gallery

Pinus rigida.JPG, Pollen cones Image:2013-05-10 09 02 57 Pitch Pine new growth and pollen cones along the Batona Trail in Brendan T. Byrne State Forest, New Jersey.jpg, New growth and pollen cones Image:Pinus rigida cone Poland.jpg, Cone and needles Image:Pitch pine cones exposed to fire.jpg, A pitch pine cone experimentally exposed to fire by Saint Michael's College scientists (Vermont; USA). The middle photograph shows the cone right after exposure to a bunsen burner flame. The right hand photograph was taken 24 hours later. File:2014-08-29 11 51 25 View north-northeast from the fire tower on Apple Pie Hill in Wharton State Forest, Tabernacle Township, New Jersey.JPG, View north from a fire tower on
Apple Pie Hill Apple Pie Hill (also known as Petty Coat Pile) is a hill in Tabernacle Township, Burlington County, New Jersey. It is tall, making it one of the highest points of the New Jersey Pine Barrens. A fire tower stands atop the summit, offering view ...
in the
New Jersey Pine Barrens The New Jersey Pine Barrens, also known as the Pinelands or simply the Pines, is the largest remaining example of the Atlantic coastal pine barrens ecosystem, stretching across more than seven counties of New Jersey. Two other large, contiguou ...
. The vast pine forest is almost entirely made up of ''Pinus rigida''. Pitch pine in Quogue (92734p).jpg, Pitch pine on Long Island in New York, USA


References

{{Authority control rigida Trees of the Eastern United States
Pinus rigida ''Pinus rigida'', the pitch pine, is a small-to-medium-sized pine. It is native to eastern North America, primarily from central Maine south to Georgia and as far west as Kentucky. It is found in environments which other species would find unsuit ...
Flora of the Appalachian Mountains Trees of the Southeastern United States Trees of the Northeastern United States Flora of Massachusetts Trees of Ontario
Pinus rigida ''Pinus rigida'', the pitch pine, is a small-to-medium-sized pine. It is native to eastern North America, primarily from central Maine south to Georgia and as far west as Kentucky. It is found in environments which other species would find unsuit ...
Pinus rigida ''Pinus rigida'', the pitch pine, is a small-to-medium-sized pine. It is native to eastern North America, primarily from central Maine south to Georgia and as far west as Kentucky. It is found in environments which other species would find unsuit ...
Garden plants of North America Plants used in bonsai
Pinus rigida ''Pinus rigida'', the pitch pine, is a small-to-medium-sized pine. It is native to eastern North America, primarily from central Maine south to Georgia and as far west as Kentucky. It is found in environments which other species would find unsuit ...
Trees of the Great Lakes region (North America) Taxa named by Philip Miller