Pisidium Javanum
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''Pisidium'' is a genus of very small or minute
freshwater clam Freshwater bivalves are one kind of freshwater mollusc, along with freshwater snails. They are bivalves that live in fresh water as opposed to salt water, which is the main habitat type for bivalves. The majority of species of bivalve molluscs l ...
s known as pill clams or pea clams, aquatic
bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
s in the family Sphaeriidae, the pea clams and fingernail clams. In some bivalve classification systems, the family Sphaeriidae is referred to as Pisidiidae, and occasionally ''Pisidium'' species are grouped in a subfamily known as Pisidiinae.


''Pisidium'' and taphonomy

In large enough quantities, the minute shells of these bivalves can affect environmental conditions, and this change in conditions can positively affect the ability of organic remains in the immediate environment to fossilize (one aspect of taphonomy). For example, in the Dinosaur Park Formation, the fossil remains of hadrosaur
eggshell An eggshell is the outer covering of a hard-shelled egg and of some forms of eggs with soft outer coats. Diversity Worm eggs Nematode eggs present a two layered structure: an external vitellin layer made of chitin that confers mechanical ...
s are rare. This is because the breakdown of tannins from the local
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant ...
vegetation caused the ancient waters to be acidic, and therefore usually eggshell fragments dissolved in the water before they had a chance to be fossilized. Hadrosaur eggshell fragments are however present in two microfossil sites in the area. Both of these sites are dominated by preserved shells of invertebrate life, primarily shells of pisidiids. The slow dissolution of these minute bivalve shells released calcium carbonate into the water, raising the water's pH high enough that it prevented the hadrosaur eggshell fragments from dissolving before they could be fossilized. Image:Pisidium moitessierianum.svg, Drawing of the right valve external view of '' Pisidium moitessierianum'' Image:Pisidium moitessierianum 3.svg, Drawing of the right valve internal view of ''Pisidium moitessierianum'' Image:Pisidium moitessierianum 2.svg, Drawing of the right valve lateral view of ''Pisidium moitessierianum'' Image:Pisidium sp.jpg, SEM image of ''Pisidium sp.''


Extant subgenera and species

Extant subgenera and species within the genus ''Pisidium'' include: Subgenus ''Euglesa'' Jenyns, 1832 *'' Pisidium casertanum'' (Poli, 1791) *'' Pisidium globulare'' Clessin, 1873 *'' Pisidium personatum'' Malm, 1855 Subgenus ''Pisidium'' Pfeiffer, 1821 *'' Pisidium amnicum'' (O. F. Müller, 1774) *'' Pisidium dilatatum'' Westerlund, 1897 Subgenus ''Cyclocalyx'' Dall, 1903 *''
Pisidium hinzi ''Euglesa hinzi'' is a species of mollusc belonging to the family Sphaeriidae. It is native to Northern Europe. Synonym: * ''Pisidium hinzi'' Kuiper, 1975 (= basionym In the scientific name of organisms, basionym or basyonym means the original ...
'' Kuiper, 1975 *'' Pisidium obtusale'' ( Lamarck, 1818) Subgenus ''Tropidocyclas'' Dall, 1903 *'' Pisidium henslowanum'' (Sheppard, 1823) *'' Pisidium lilljeborgii'' Clessin, 1886 *'' Pisidium supinum'' A. Schmidt, 1851 *''
Pisidium waldeni ''Euglesa waldeni'' is a species of mollusc belonging to the family Sphaeriidae. It is native to Northern Europe. Synonym: * ''Pisidium waldeni'' Kuiper, 1975 (= basionym In the scientific name of organisms, basionym or basyonym means the orig ...
'' Kuiper, 1975 Subgenus ''Hiberneuglesa'' Starobogatov, 1983 *'' Pisidium hibernicum'' Westerlund, 1894 Subgenus ''Cingulipisidium'' Pirogov & Starobogatov, 1974 *'' Pisidium crassum'' Stelfox, 1918 *''
Pisidium milium ''Pisidium milium'' is a species of very small freshwater bivalve in the family Sphaeriidae, the fingernail clams and pea clams. Description The 3.0- to 4.5-mm shell is tumid (swollen). It is more rectangular than other ''Pisidium'' specie ...
'' Held, 1836 *''
Pisidium nitidum ''Pisidium nitidum'', the shining pea clam, is a species of minute freshwater clam, an aquatic bivalve mollusc in the family Sphaeriidae, the pea clams and fingernail clams. Description The 3 – 4 mm. shell is slightly tumid (swollen). ...
'' Jenyns, 1832 *'' Pisidium pseudosphaerium'' Favre, 1927 Subgenus ''Pseudeupera'' Germain, 1909 *''
Pisidium pulchellum ''Pisidium pulchellum'' , the iridescent pea mussel, is a minute species of pea clam, a freshwater bivalve in the family Sphaeriidae. Description The 3.5–4 mm shell is tumid (swollen), oblique-oval shell. The umbos are broad well and w ...
'' Jenyns, 1832 *'' Pisidium subtruncatum'' Malm, 1855 Subgenus ''Neopisidium'' Odhner, 1921 *''
Pisidium conventus ''Pisidium conventus'', the arctic-alpine pea clam, is a species of freshwater bivalve from the family Sphaeriidae. Description The small (2.2–2.8 mm.) shell is oval in shape and thin and fragile. It is broad and flattened (laterally com ...
'' Clessin, 1867 Subgenus ''Odhneripisidium'' Kuiper, 1962 *'' Pisidium moitessierianum'' Paladilhe, 1866 *''
Pisidium tenuilineatum ''Pisidium tenuilineatum'', the fine-lined pea mussel, is a species of very small freshwater bivalve in the family Sphaeriidae. Distribution The species is native to Europe. * British Isles – listed in List of endangered species in the Britis ...
'' Stelfox, 1918 Subgenus ''Afropisidium'' Kuiper, 1962 *'' Pisidium giraudi'' Bourguignat, 1885 *''
Pisidium hodgkini ''Pisidium hodgkini'' is a freshwater bivalve of the family Sphaeriidae, which is found in New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North ...
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, 1905) *''
Pisidium pirothi ''Afropisidium pirothi'' is a species of bivalve belonging to the family Sphaeriidae. The species is found in Africa. References

Sphaeriidae Bivalves described in 1881 Bivalves of Africa {{Sphaeriidae-stub ...
'' Jickeli, 1881 Subgenus unknown *''
Pisidium artifex ''Pisidium artifex'' is a species of freshwater clam in the family Sphaeriidae. It is endemic to Kenya, where it is known only from Mount Kenya Mount Kenya (Kikuyu: ''Kĩrĩnyaga'', Kamba, ''Ki Nyaa'') is the highest mountain in Kenya and t ...
'' Kuiper, 1960 *'' Pisidium stewarti'' Preston, 1909 Subgenus ' *''
Pisidium annandalei ''Pisidium'' is a genera, genus of very small or minute freshwater clams known as pill clams or pea clams, Aquatic animal, aquatic bivalve molluscs in the family (biology), family Sphaeriidae, the pea clams and fingernail clams. In some bivalv ...
'' Prashad, 1925 *''
Pisidium edlaueri ''Pisidium'' is a genus of very small or minute freshwater clams known as pill clams or pea clams, aquatic bivalve molluscs in the family Sphaeriidae, the pea clams and fingernail clams. In some bivalve classification systems, the family Spha ...
'' Kuiper, 1960 *'' Pisidium javanum'' van Benthem Jutting, 1931 *''
Pisidium maasseni ''Euglesa maasseni'' is a species of bivalve belonging to the family Sphaeriidae Sphaeriidae is a family of small to minute freshwater bivalve molluscs in the order Sphaeriida. In the US, they are commonly known as pea clams or fingernail cl ...
'' Kuiper, 1987 *''
Pisidium punctiferum ''Pisidium punctiferum'', the striate peaclam, is a freshwater bivalve of the family (biology), family Sphaeriidae. Distribution The type locality is a pond at Saint Ann's River, Trinidad and Tobago, Saint Ann's River, near Port of Spain, Tri ...
'' (Guppy, 1867) *''
Pisidium raddei ''Pisidium'' is a genus of very small or minute freshwater clams known as pill clams or pea clams, aquatic bivalve molluscs in the family Sphaeriidae, the pea clams and fingernail clams. In some bivalve classification systems, the family Spha ...
'' Dybowski, 1902 * undescribed ''Pisidium'' species from Africa (Kuiper, in prep., awaiting additional records)Kuiper J.G.J. (2009). "Fossil records of Palaearctic ''Pisidium'' species in tropical Africa". Zoologische Mededelingen 83(10): 593-594
HTM
.


Footnotes


References

* Tanke, D.H. and Brett-Surman, M.K. 2001. Evidence of Hatchling and Nestling-Size Hadrosaurs (Reptilia:Ornithischia) from Dinosaur Provincial Park (Dinosaur Park Formation: Campanian), Alberta, Canada. pp. 206–218. In: Mesozoic Vertebrate Life—New Research Inspired by the Paleontology of Philip J. Currie. Edited by D.H. Tanke and K. Carpenter. Indiana University Press: Bloomington. xviii + 577 pp.


External links


''Pisidium''
keyed, described and illustrated in ''Danmarks Fauna'' (Georg Mandahl-Barth)
''Pisidium'' images
at Consortium for the Barcode of Life * https://web.archive.org/web/20141128091542/http://sunsite.ualberta.ca/Projects/Aquatic_Invertebrates/?Page=20 {{Taxonbar, from=Q600521 Bivalve genera