''Pinus peregrinus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...

in the family

Pinaceae The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly kn ...

known from

Clarkforkian The Clarkforkian North American Stage, on the geologic timescale, is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 56,800,000 to 55,400,000 years BP lasting . Considered t ...

age

Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s found in western

North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the Native Americans in the United States, indigenous Dakota people, Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north a ...

, USA. The species was described from three wing seed specimens found at localities 14051a and 14083 in the Bear Den member of the

Golden Valley Formation The Golden Valley Formation is a stratigraphic unit of Late Paleocene to Early Eocene age in the Williston Basin of North Dakota.Hickey, 1977 It is present in western North Dakota and was named for the city of Golden Valley by W.E. Benson and W.M ...

with associated needles from the Heart River Bluffs area, site 14051a, provisionally assigned to the species also. The Bear Den member outcrops at a number of sites in western

, and is designated the type locality. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

specimen, number ''PU 20091'', is preserved in the

Princeton University Princeton University is a private university, private research university in Princeton, New Jersey. Founded in 1746 in Elizabeth, New Jersey, Elizabeth as the College of New Jersey, Princeton is the List of Colonial Colleges, fourth-oldest ins ...

collections, and

paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). O ...

specimens are in the

National Museum of Natural History The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7 ...

collections of the

Smithsonian Institution The Smithsonian Institution ( ), or simply the Smithsonian, is a group of museums and education and research centers, the largest such complex in the world, created by the U.S. government "for the increase and diffusion of knowledge". Founded ...

. The specimens were studied by

paleobotanist Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeogr ...

Leo J. Hickey Leo or Léo may refer to: Acronyms * Law enforcement officer * Law enforcement organisation * ''Louisville Eccentric Observer'', a free weekly newspaper in Louisville, Kentucky * Michigan Department of Labor and Economic Opportunity Arts an ...

of the Yale University Geology Department. Dr Hickey published the 1977

type description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have be ...

for ''P. peregrinus'' in the

Geological Society of America The Geological Society of America (GSA) is a nonprofit organization dedicated to the advancement of the geosciences. History The society was founded in Ithaca, New York, in 1888 by Alexander Winchell, John J. Stevenson, Charles H. Hitch ...

memoir 150, ''Stratigraphy and Paleobotany of the Golden Valley Formation (Early Tertiary) of Western North Dakota''. Dr Hickey chose the specific name ''peregrinus'', which is

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

meaning "stranger" or "newcomer" noting the species to be the first

megafossil Macrofossils, also known as megafossils, are preserved organic remains large enough to be visible without a microscope. The term ''macrofossil'' stands in opposition to the term microfossil. Microfossils, by contrast, require substantial magnific ...

record for the

pine family The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, larches, pines and spruces. The family is included in the order Pinales, ...

to be described from

Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...

and

Great Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, an ...

Paleocene rocks. The winged seeds of ''Pinus peregrinus'' are between long and have a nutlet located in the base of the wing to one side of the wing axis. The small, long nutlet is three sided and generally deltoid to ovoid in shape. The wing possesses a thickened inside margin that is straight, while the thin outside margin arches from nutlet to distal tip of the wing. The wing membrane is slightly striated parallel to the wing margins and curving towards the outside margin near the tip. This combination of characters is specific to the genus ''

Pinus A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden ...

'', and the seeds are most similar to the modern ''

Pinus resinosa ''Pinus resinosa'', known as red pine (also Norway pine in Minnesota), is a pine native to North America. Description Red pine is a coniferous evergreen tree characterized by tall, straight growth. It usually ranges from in height and in trun ...

'' and ''

Pinus tropicalis ''Pinus tropicalis'', the tropical pine, is a pine tree endemic to the western highlands of the island of Cuba. See also *Cuban pine forests The Cuban pine forests are a tropical coniferous forest ecoregion on the Caribbean islands of Cuba an ...

''. The needles provisionally included in ''P. peregrinus'' can reach up to long with a distinct midrib.

References

{{Taxonbar, from=Q7196567 peregrinus Plants described in 1977 Paleocene plants Extinct flora of North America Prehistoric trees