''Pinus driftwoodensis'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...

in the

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...

family solely known from

early Eocene In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian i ...

sediments exposed in south central

British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...

. The species was described from an isolated

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

ovulate cone associated with a series of wood, needles and pollen cones in

chert Chert () is a hard, fine-grained sedimentary rock composed of microcrystalline or cryptocrystalline quartz, the mineral form of silicon dioxide (SiO2). Chert is characteristically of biological origin, but may also occur inorganically as a prec ...

. ''P. driftwoodensis'' was the eighth pine species to be described from a

permineralized Permineralization is a process of fossilization of bones and tissues in which mineral deposits form internal casts of organisms. Carried by water, these minerals fill the spaces within organic tissue. Because of the nature of the casts, perminera ...

ovulate cone and the second from the Okanagan highlands (after '' Pinus arnoldii'').

History and classification

''Pinus driftwoodensis'' has been identified from a single location, the type locality, at the Driftwood Canyon exposure of Ootsa Lake Group sediments near

Smithers, British Columbia Smithers is a town in northwestern British Columbia, approximately halfway between Prince George and Prince Rupert. With a population of 5,351 in 2016, Smithers provides service coverage for most of the Bulkley Valley. History Region First Nation ...

in the

Driftwood Canyon Provincial Park Driftwood Canyon Provincial Park is a provincial park in British Columbia, Canada. Driftwood Canyon Provincial Park covers 23 ha of the Bulkley River Valley, on the east side of Driftwood Creek, a tributary of the Bulkley River, 10 km northe ...

. The Driftwood Canyon location is currently considered to be Early Eocene in age, based on Uranium-lead

radiometric dating Radiometric dating, radioactive dating or radioisotope dating is a technique which is used to date materials such as rocks or carbon, in which trace radioactive impurities were selectively incorporated when they were formed. The method compares t ...

which yielded an age of 51.77 ± 0.34

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

. Sediments at Driftwood canyon are interpreted as preserving a lacustrian upland flora and fauna with the chert blocks being found in a small coal and chert lens interbedded with Oostsa Lake group shales. The species was described from a single

type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...

, the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

specimen S5446, which is currently preserved in the

University of Alberta The University of Alberta, also known as U of A or UAlberta, is a public research university located in Edmonton, Alberta, Canada. It was founded in 1908 by Alexander Cameron Rutherford,"A Gentleman of Strathcona – Alexander Cameron Rutherfor ...

paleobotanical collections in

Edmonton, Alberta Edmonton ( ) is the capital city of the Canadian province of Alberta. Edmonton is situated on the North Saskatchewan River and is the centre of the Edmonton Metropolitan Region, which is surrounded by Alberta's central region. The city anchor ...

. Specimen S5446 is a block of chert which preserves the ovulate cone, two types of needle foliage, several branch segments, and several small pollen cone. The specimen were studied by

paleobotanist Paleobotany, which is also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeogr ...

Ruth A. Stockey of the

. She published her 1983

type description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have be ...

for ''P. driftwoodensis'' in the journal '' Botanical Gazette''. In her type description she did not note the

etymology Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the Phonological chan ...

for the

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''driftwoodensis''.

Description

The ovulate cone of ''Pinus driftwoodensis'' is rounded with a pointed tip with an overall length of by wide and was missing the base before preservation occurred. Based on the interrolling of the cone scales on the basal preserved area of the cone indicates the missing section to be no more the . ''P. driftwoodensis'' notably differs from the

coeval {{Short pages monitor