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The pink-headed duck (''Rhodonessa caryophyllacea'') is a large
diving duck The diving ducks, commonly called pochards or scaups, are a category of duck which feed by diving beneath the surface of the water. They are part of Anatidae, the diverse and very large family that includes ducks, geese, and swans. The diving d ...
that was once found in parts of the Gangetic plains of India, parts of Maharashtra, Bangladesh and in the riverine swamps of Myanmar but feared extinct since the 1950s. Numerous searches have failed to provide any proof of continued existence. It has been suggested that it may exist in the inaccessible swamp regions of northern Myanmar and some sight reports from that region have led to its status being declared as "Critically Endangered" rather than extinct. The genus placement has been disputed and while some have suggested that it is close to the
red-crested pochard The red-crested pochard (''Netta rufina'') is a large diving duck. The scientific name is derived from Greek ''Netta'' "duck", and Latin ''rufina'', "golden-red" (from ''rufus'', "ruddy"). Its breeding habitat is lowland marshes and lakes in sout ...
(''
Netta rufina The red-crested pochard (''Netta rufina'') is a large diving duck. The scientific name is derived from Greek ''Netta'' "duck", and Latin ''rufina'', "golden-red" (from ''rufus'', "ruddy"). Its breeding habitat is lowland marshes and lakes in sout ...
''), others have placed it in a separate genus of its own. It is unique in the pink colouration of the head combined with a dark body. A prominent wing patch and the long slender neck are features shared with the common
Indian spot-billed duck The Indian spot-billed duck (''Anas poecilorhyncha'') is a large dabbling duck that is a non-migratory breeding duck throughout freshwater wetlands in the Indian subcontinent. The name is derived from the red spot at the base of the bill that is ...
. The eggs have also been held as particularly peculiar in being nearly spherical.


Description

The male pink-headed duck is unmistakable when a good view is had. Both sexes are 41–43 cm and long-billed with long necks and peaked heads. The male has a pink bill, head and neck while the female has a pale pinkish head and neck with a paler bill. The black of the body extends as a narrow strip on the front of the neck. Wings have a leading white edge. In flight it would not contrast as much as the syntopic white-winged duck. Wing does not have the dark trailing edge of the
red-crested pochard The red-crested pochard (''Netta rufina'') is a large diving duck. The scientific name is derived from Greek ''Netta'' "duck", and Latin ''rufina'', "golden-red" (from ''rufus'', "ruddy"). Its breeding habitat is lowland marshes and lakes in sout ...
. Confusion with male red-crested pochards stems mainly from observations of swimming birds, as the latter species also has a conspicuous red head (although the color is actually very different from the pink-headed duck).
Indian spot-billed duck The Indian spot-billed duck (''Anas poecilorhyncha'') is a large dabbling duck that is a non-migratory breeding duck throughout freshwater wetlands in the Indian subcontinent. The name is derived from the red spot at the base of the bill that is ...
s, on the other hand, can look similar to female pink-headed ducks when in flight and seen from a distance, and if seen from behind, they could be mistaken for males too. The upper side of the wing is distinguishing, with dark green secondaries (speculum) and prominent white tertiaries in the spot-billed duck and a pinkish-beige speculum, much lighter than its surroundings, in the pink-headed duck. If the upper part of the wings cannot be reliably seen, they are all but indistinguishable except to expert observers in good visibility conditions. Young birds had a nearly whitish head without a trace of pink and a mellow two note call ''wugh-ah'' has been attributed to the species. Its breeding habitat is lowland
marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
es and pools in tall-grass jungle. The nest is built amongst grass. The eggs, six or seven in a clutch, are very spherical and creamy white. The eggs measure 1.71 to 1.82 inches long and 1.61 to 1.7 inches wide. They were believed to have been non-migratory and found singly or in pairs and very rarely in small groups. Pink-headed ducks are believed to have eaten water plants and molluscs. Like ''
Netta ''Netta'' is a genus of diving ducks. The name is derived from Greek ''Netta'' "duck". Unlike other diving ducks, the ''Netta'' species are reluctant to dive, and feed more like dabbling ducks. These are gregarious ducks, mainly found on fresh ...
'' species, they typically up-ended or dabbled for food and did not dive like a pochard.


Distribution

Allan Octavian Hume Allan Octavian Hume, CB ICS (4 June 1829 – 31 July 1912) was a British civil servant, political reformer, ornithologist and botanist who worked in British India. He was the founder of the Indian National Congress. A notable ornithologist, Hum ...
and Stuart Baker noted that the stronghold of the species was north of the Ganges and west of the Brahmaputra, mainly in Maldah, Purnea, Madhubani and Purulia districts of present-day Bihar. It was said to be commoner in Singhbum. Hume collected a specimen in Manipur which he noted was very rare, hiding among dense reeds in
Loktak Lake , image = , caption = Different scenes of the Loktak lake of Manipur , alt = View of Loktak Lake and Phumdis , image_bathymetry = , caption_bathymetry = , location = Manipur , coords ...
.
Edward Blyth Edward Blyth (23 December 1810 – 27 December 1873) was an English zoologist who worked for most of his life in India as a curator of zoology at the museum of the Asiatic Society of India in Calcutta. Blyth was born in London in 1810. In 1841 ...
claimed that it was found in the Rakhine state of Burma.
Brian Houghton Hodgson Brian Houghton Hodgson (1 February 1800 or more likely 1801 – 23 May 1894) was a pioneer naturalist and ethnologist working in India and Nepal where he was a British Resident. He described numerous species of birds and mammals from the Himala ...
obtained specimens from Nepal. A few records were also noted from Delhi, Sindh and Punjab. The populations (possibly) undertook local seasonal movements, resulting in scattered historic records as far as Punjab, Maharahtra and Andhra Pradesh. Thus, resulting in rare sightings of the species in Maharashtra. Birds were also reported from the Oudh region some from very close to Lucknow. Specimens were shot at Najafgarh lake in the Delhi district. Jerdon obtained specimens of the bird from further south although he did not personally observe any in the wild until he visited Bengal.


Taxonomy and systematics

The pink-headed duck was described by John Latham in 1790 under the genus ''Anas''. In describing the species, it is possible that he made of use of a painting in the collection of Lady Impey, wife of Sir
Elijah Impey Sir Elijah Impey (13 June 17321 October 1809) was a British judge, the first chief justice of the Supreme Court of Judicature at Fort William in Bengal, Chief Justice of the Sadr Diwani Adalat and MP for New Romney. Life He was born the young ...
who was Chief Justice of court in Calcutta from 1774 to 1783. Mary Impey maintained a menagerie in Calcutta and commissioned Indian artists such as Bhawani Das from Patna to illustrate animals in the collection. The Impeys moved to England, and after the death of her husband, she sold these paintings at auction in 1810. Some of them were acquired by the
13th Earl of Derby Edward Smith-Stanley, 13th Earl of Derby (21 April 1775 – 30 June 1851), KG, of Knowsley Hall in Lancashire (styled Lord Stanley from 1776 to 1832, known as Baron Stanley of Bickerstaffe from 1832-4), was a politician, peer, landowner, bui ...
. The genus ''Rhodonessa'' was originally created for this species alone.
Jean Delacour Jean may refer to: People * Jean (female given name) * Jean (male given name) * Jean (surname) Fictional characters * Jean Grey, a Marvel Comics character * Jean Valjean, fictional character in novel ''Les Misérables'' and its adaptations * Jea ...
and
Ernst Mayr Ernst Walter Mayr (; 5 July 1904 – 3 February 2005) was one of the 20th century's leading evolutionary biologists. He was also a renowned Taxonomy (biology), taxonomist, tropical explorer, ornithologist, Philosophy of biology, philosopher o ...
, in their 1945 revision of the family Anatidae considered it a somewhat abnormal member of the ''Anatini'' (or river-ducks) group because the hind toe is slightly lobed, display behaviour and the tendency to feed at the surface. The birds were observed in European aviaries and although they never bred, the males displayed often and this involved puffing the neck feathers, lowering the neck to rest on the back and then stretching up the neck while producing a wheezy whistle like a
mallard The mallard () or wild duck (''Anas platyrhynchos'') is a dabbling duck that breeds throughout the temperate and subtropical Americas, Eurasia, and North Africa, and has been introduced to New Zealand, Australia, Peru, Brazil, Uruguay, Arge ...
. A study of its tracheal anatomy by
Alfred Henry Garrod Alfred Henry Garrod FRS (May 18, 1846 – October 17, 1879) was an English vertebrate zoologist. Garrod was born in London, the eldest son of Sir Alfred Baring Garrod (1819–1907), a physician at King's College Hospital, who discovered ...
in 1875 suggested that it had a "slight fusiform dilatation" in the anterior syringeal region. The "bulba ossea" at the lower part of the male syrinx is peculiar in being swollen. The colour pattern has also been considered unique, lacking any of the metallic colours on the secondaries that are characteristic of the Anatini. The other unique feature being the somewhat large and nearly spherical shape of the eggs. All of these features supporting the retention of the species in a separate genus. Such mid-tracheal swellings were found only in
Mergini The sea ducks (''Mergini'') are a tribe of the duck subfamily of birds, the Anatinae. The taxonomy of this group is incomplete. Some authorities separate the group as a subfamily, while others remove some genera. Most species within the group sp ...
and
Aythyini The diving ducks, commonly called pochards or scaups, are a category of duck which feed by diving beneath the surface of the water. They are part of Anatidae, the diverse and very large family that includes ducks, geese, and swans. The diving d ...
and is extremely rare in the genus ''
Anas ''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus wa ...
''. This tracheal bulla is rounded in ''Anas'' but angular with fenestrae in ''
Netta ''Netta'' is a genus of diving ducks. The name is derived from Greek ''Netta'' "duck". Unlike other diving ducks, the ''Netta'' species are reluctant to dive, and feed more like dabbling ducks. These are gregarious ducks, mainly found on fresh ...
'' and ''
Aythya ''Aythya'' is a genus of diving ducks. It has twelve described species. The name ''Aythya'' comes from the Ancient Greek word (), which may have referred to a sea-dwelling duck or an auklet. ''Aythya shihuibas'' was described from the Late Mio ...
''. Johnsgard considered ''
Marmaronetta The marbled duck, or marbled teal (''Marmaronetta angustirostris'') is a medium-sized species of duck from southern Europe, northern Africa, and western and central Asia. The scientific name, ''Marmaronetta angustirostris'', comes from the Greek ...
'' and ''Rhodonessa'' as intermediate in form. Based on the available morphological and behavioural evidence, especially the structure of the
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
and the structure of
tracheal The trachea, also known as the windpipe, is a cartilaginous tube that connects the larynx to the bronchi of the lungs, allowing the passage of air, and so is present in almost all air-breathing animals with lungs. The trachea extends from the la ...
rings,
Sidney Dillon Ripley Sidney Dillon Ripley II (September 20, 1913 – March 12, 2001) was an American ornithologist and wildlife conservationist. He served as secretary of the Smithsonian Institution for 20 years, from 1964 to 1984, leading the institution through ...
suggested that it was undoubtedly in the Aythyini. A study found that ''Rhodonessa'' was closely allied to the
red-crested pochard The red-crested pochard (''Netta rufina'') is a large diving duck. The scientific name is derived from Greek ''Netta'' "duck", and Latin ''rufina'', "golden-red" (from ''rufus'', "ruddy"). Its breeding habitat is lowland marshes and lakes in sout ...
(''Netta rufina'') suggesting that the two species be placed in the same genus. ''Rhodonessa'' was described prior to ''Netta'' which would then make ''Rhodonessa rufina'' the name of choice, however these changes have not been widely accepted. The pink colour is derived from a carotenoid pigment which is unusual among ducks and known only from a few other species such as the
pink-eared duck The pink-eared duck (''Malacorhynchus membranaceus'') is a species of duck found in Australia. It has a large spatulate bill like the Australasian shoveler, but is smaller at 38–40 cm length. Its brown back and crown, black and white barr ...
which are not closely related.


Status

This
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form t ...
formerly occurred in eastern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
and northern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, but is now probably
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
. It was always rare, and the last confirmed sighting, by C. M. Inglis, was from Bhagownie, Darbhangha District, in June 1935, with reports from India persisting until the early 1960s. These include reports from
Monghyr Munger, formerly spelt as Monghyr, is a twin city and a Municipal Corporation situated in the Indian state of Bihar. It is the administrative headquarters of Munger district and Munger Division. Munger was one of the major cities in Eastern In ...
and from near
Shimla Shimla (; ; also known as Simla, List of renamed Indian cities and states#Himachal Pradesh, the official name until 1972) is the capital and the largest city of the States and union territories of India, northern Indian state of Himachal Prade ...
.
Sidney Dillon Ripley Sidney Dillon Ripley II (September 20, 1913 – March 12, 2001) was an American ornithologist and wildlife conservationist. He served as secretary of the Smithsonian Institution for 20 years, from 1964 to 1984, leading the institution through ...
considered it likely extinct in 1950. In 1988, Rory Nugent, an American birder, and Shankar Barua of
Delhi Delhi, officially the National Capital Territory (NCT) of Delhi, is a city and a union territory of India containing New Delhi, the capital of India. Straddling the Yamuna river, primarily its western or right bank, Delhi shares borders w ...
, reported spotting the elusive bird on the banks of the
Brahmaputra The Brahmaputra is a trans-boundary river which flows through Tibet, northeast India, and Bangladesh. It is also known as the Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, Luit in Assamese, and Jamuna River in Bangla. It ...
. The pair started their quest for the bird at Saikhoa ghat on the north-eastern end of the river on the Indian side of the border. After 29 days of sailing, Nugent said that he saw the pink-headed duck amidst a flock of other waterbirds. However, Nugent and Barua's claimed sighting has not been widely accepted. Reports of pink-headed ducks after the 1960s have been received from the largely unexplored
Mali Hka The Mali River (''Mali Hka'') is a river that originates in the hills of Kachin State, in the northern border of Myanmar. It flows approximately 320 km, when it meets with the N'Mai River and forms the Ayeyarwady River. History Constructi ...
and Chindwin Myit drainages in Northern Myanmar. While the area is not very well surveyed by scientists, searches have been inconclusive and confusion with the
red-crested pochard The red-crested pochard (''Netta rufina'') is a large diving duck. The scientific name is derived from Greek ''Netta'' "duck", and Latin ''rufina'', "golden-red" (from ''rufus'', "ruddy"). Its breeding habitat is lowland marshes and lakes in sout ...
and the
Indian spot-billed duck The Indian spot-billed duck (''Anas poecilorhyncha'') is a large dabbling duck that is a non-migratory breeding duck throughout freshwater wetlands in the Indian subcontinent. The name is derived from the red spot at the base of the bill that is ...
has been a common source of supposed pink-headed duck sightings. A report on a survey in the Hu Kaung valley in November 2003 concluded that there is sufficient reason to believe that pink-headed ducks may still exist in Northern Myanmar's
Kachin State Kachin State ( my, ကချင်ပြည်နယ်; Kachin: ), also known by the endonym Kachinland, is the northernmost state of Myanmar. It is bordered by China to the north and east (Tibet and Yunnan, specifically and respectively); Sh ...
, but a thorough survey of the Nat Kaung river between
Kamaing Kamaing ( my, ကာမိုင်းမြို့; also Kamine) is a jade-mining town in the Kachin State of the northernmost part of the Republic of the Union of Myanmar. It is the birthplace of journalist Edward Michael Law-Yone Edward Mi ...
and Shadusup in October 2005 failed to find this species; a number of interesting ducks were observed, but they turned out to be
Indian spot-billed duck The Indian spot-billed duck (''Anas poecilorhyncha'') is a large dabbling duck that is a non-migratory breeding duck throughout freshwater wetlands in the Indian subcontinent. The name is derived from the red spot at the base of the bill that is ...
s or
white-winged duck The white-winged duck or white-winged wood duck (''Asarcornis scutulata'') is a large species of duck, formerly placed in the genus ''Cairina'' with the Muscovy duck (''Cairina moschata'') and allied with the dabbling ducks. However, mtDNA cyt ...
s. Suggestions have been made that it may be nocturnal. In 2017, an expedition to find the species by Global Wildlife Conservation also failed, with evidence indicating that the biodiversity in the general area around
Indawgyi Lake Indawgyi Lake ( ) is one of the largest inland lakes in Southeast Asia. It is located in Mohnyin Township in the Kachin State of Myanmar. The lake measures east to west, and north to south. There are over 20 villages around the lake. The predomi ...
and its surrounding areas was heavily declining due to habitat degradation. Anecdotes from residents in the area, however, indicate that the bird may have lived in the area far more recently than the last confirmed report from 1910, possibly as recently as 2010. One resident stated that a pink-headed duck was sighted in 1998, associating with a flock of
gadwall The gadwall (''Mareca strepera'') is a common and widespread dabbling duck in the family Anatidae. Taxonomy The gadwall was first described by Carl Linnaeus in his landmark 1758 10th edition of ''Systema Naturae''. DNA studies have shown that ...
and pintail. Another, more dubious report stated that shortly after a failed expedition in the area by
Birdlife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
ended, a local hunter caught a live male and a female or juvenile pink-headed duck, and contacted Myanmar's Biodiversity and Nature Conservation Association to sell it to them for a fee. The offer was declined, and the hunter killed both ducks. Another hunter recalled that when the habitat was in good condition, pink-headed ducks were regulars in the area, possibly up to 2014. They were apparently most common during February, and he also could mimic their possible calls, though it is unknown whether these calls were truly by pink-headed ducks. The hunter also said that there were large, impassable ponds in the wetland's center that may still hold pink-headed ducks, but these could only be accessed with a drone, which are banned in the region. The reason for its disappearance was probably habitat destruction. It is not known why it was always considered rare, but the rarity is believed to be genuine (and not an artefact of insufficient fieldwork) as its erstwhile habitat was frequently scoured by hunters in Colonial times. The pink-headed duck was much sought after by hunters and later as an ornamental bird, mainly because of its unusual plumage. Like most diving ducks, it was not considered good eating, which should facilitate the survival of any remnant birds. The last specimen was obtained in 1935 in
Darbhanga Darbhanga is the fifth-largest city and municipal corporation in the Indian state of Bihar situated centrally in Mithila region. Darbhanga is the headquarters of the Darbhanga district and the Darbhanga division. It was the seat of the erstw ...
,
Andhra Pradesh Andhra Pradesh (, abbr. AP) is a state in the south-eastern coastal region of India. It is the seventh-largest state by area covering an area of and tenth-most populous state with 49,386,799 inhabitants. It is bordered by Telangana to the ...
, India by C. M. Inglis. Some birds were also kept in the aviaries of
Jean Théodore Delacour Jean Théodore Delacour (26 September 1890 – 5 November 1985) was a French ornithologist and aviculturist. He later became American. He was renowned for not only discovering but also rearing some of the rarest birds in the world. He establish ...
in
Clères Clères () is a commune in the Seine-Maritime department in the Normandy region in north-western France. Geography A farming and forestry market town situated by the banks of the Clérette river, in the Pays de Caux, some northeast of Rouen, ...
(
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
) and Alfred Ezra at
Foxwarren Park Foxwarren Park, at Wisley in Surrey, is a Victorian country house and estate. On sandstone Ockham and Wisley Commons, it was designed in 1860 by the railway architect Frederick Barnes for brewing magnate and MP, Charles Buxton. It is a Grade ...
(
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
) where the last known birds lived in captivity. The only known photographs of the species were taken here and include one of a pair taken around 1925 by
David Seth-Smith David Seth-Smith FZS, MBOU (9 March 1875 – 30 October 1963) was a British zoologist, wildlife artist, nature broadcaster and author. His career included spells as Curator of Mammals and Birds for the Zoological Society of London and edit ...
.


References


Other sources

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External links


BirdLife International Species Factsheet
{{Authority control
pink-headed duck The pink-headed duck (''Rhodonessa caryophyllacea'') is a large diving duck that was once found in parts of the Gangetic plains of India, parts of Maharashtra, Bangladesh and in the riverine swamps of Myanmar but feared extinct since the 1950s. ...
pink-headed duck The pink-headed duck (''Rhodonessa caryophyllacea'') is a large diving duck that was once found in parts of the Gangetic plains of India, parts of Maharashtra, Bangladesh and in the riverine swamps of Myanmar but feared extinct since the 1950s. ...
Birds of India Birds of Myanmar Birds of Bangladesh
pink-headed duck The pink-headed duck (''Rhodonessa caryophyllacea'') is a large diving duck that was once found in parts of the Gangetic plains of India, parts of Maharashtra, Bangladesh and in the riverine swamps of Myanmar but feared extinct since the 1950s. ...
pink-headed duck The pink-headed duck (''Rhodonessa caryophyllacea'') is a large diving duck that was once found in parts of the Gangetic plains of India, parts of Maharashtra, Bangladesh and in the riverine swamps of Myanmar but feared extinct since the 1950s. ...
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