The pine grosbeak (''Pinicola enucleator'') is a large member of the true
finch
The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
family
Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
,
Fringillidae
The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
. It is the only species in the genus ''Pinicola''. It is found in
conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous woods across
Alaska
Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
, the western mountains of the
United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
,
Canada
Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
, and in
subarctic
The subarctic zone is a region in the Northern Hemisphere immediately south of the true Arctic, north of humid continental regions and covering much of Alaska, Canada, Iceland, the north of Scandinavia, Siberia, and the Cairngorms. Generally, ...
Fennoscandia
__NOTOC__
Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and no, Fennoskandia, nocat=1; russian: Фенноскандия, Fennoskandiya) or the Fennoscandian Peninsula is the geographical peninsula in Europe, which includes ...
and across the
Palearctic
The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa.
The realm consists of several bioregions: the Euro-Sibe ...
to
Siberia
Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
. The species is a
frugivore
A frugivore is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance an ...
, especially in winter, favoring small fruits, such as
rowan
The rowans ( or ) or mountain-ashes are shrubs or trees in the genus ''Sorbus
''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' (''s.l.'') are commonly known as whitebeam, r ...
s (mountain-ashes in the New World). With fruit-crop abundance varying from year to year, pine grosbeak is one of many subarctic-resident bird species that exhibit irruptive behavior. In
irruption years, individuals can move long distances in search of suitable food supplies, bringing them farther south and/or downslope than is typical of years with large fruit crops.
Taxonomy
The pine grosbeak was
formally described in 1758 by the Swedish naturalist
Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in the
tenth edition of his ''
Systema Naturae
' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' under the
binomial name
In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Loxia enucleator''. The
type locality is Sweden. It is now the only species placed in the genus ''Pinicola'' that was erected in 1808 by the French ornithologist
Louis Jean Pierre Vieillot
Louis Pierre Vieillot (10 May 1748, Yvetot – 24 August 1830, Sotteville-lès-Rouen) was a French ornithologist.
Vieillot is the author of the first scientific descriptions and Linnaean names of a number of birds, including species he collecte ...
.
The generic name ''Pinicola'' combines the Latin ''pinus'' meaning "pine tree" and ''colere'' meaning "to dwell"; the specific ''enucleator'' is from the Latin ''enucleare'' meaning "to remove the kernel".
The pine grosbeak is a
sister taxon
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree.
Definition
The expression is most easily illustrated by a cladogram:
Taxon A and t ...
to the bullfinches of the genus ''
Pyrrhula
Pyrrhula is a small genus of passerine birds, commonly called bullfinches, belonging to the finch family (Fringillidae). The genus has a Palearctic distribution; almost all species occur in Asia, with two species exclusively in the Himalayas and ...
''.
The
crimson-browed finch
The crimson-browed finch (''Carpodacus subhimachalus'') is a true finch species (family Fringillidae). It is found in Bhutan, China, India, Myanmar, and Nepal. Its natural habitats are temperate forests and temperate shrubland.
This is a large f ...
(''Carpodacus subhimachalus'') was previously placed in the genus ''Pinicola''. It was moved to the rosefinch genus ''
Carpodacus
The rosefinches are a genus, ''Carpodacus'', of passerine birds in the finch family Fringillidae. Most are called "rosefinches" and as the word implies, have various shades of red in their plumage. The common rosefinch is frequently called the ...
'' based on results from phylogenetic studies of
mitochondrial
A mitochondrion (; ) is an organelle found in the cells of most Eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is use ...
and nuclear DNA sequences that were published in 2012 and 2013.
[
Eight ]subspecies
In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised:[
* ''P. e. enucleator'' (Linnaeus, 1758) – Scandinavia to central Siberia
* ''P. e. kamtschatkensis'' ( Dybowski, 1883) – northeastern Siberia
* ''P. e. sakhalinensis'' ]Buturlin Buturlin, feminine: Buturlina (russian: Бутурли́н, Бутурлина) is a Russian surname of a Russian noble . Notable people with this surname include:
* Alexander Buturlin (1694–1767), a Russian general
* Anna Artemevna Buturlina ( ...
, 1915 – Sakhalin
Sakhalin ( rus, Сахали́н, r=Sakhalín, p=səxɐˈlʲin; ja, 樺太 ''Karafuto''; zh, c=, p=Kùyèdǎo, s=库页岛, t=庫頁島; Manchu: ᠰᠠᡥᠠᠯᡳᠶᠠᠨ, ''Sahaliyan''; Orok: Бугата на̄, ''Bugata nā''; Nivkh: ...
and the Kuril Islands
The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese: or ) are a volcanic archipelago currently administered as part of Sakhalin Oblast in the ...
, northern Japan
* ''P. e. flammula'' Homeyer, 1880 – coastal southern Alaska and western Canada
* ''P. e. carlottae'' Brooks, AC, 1922 – the Queen Charlotte Islands
Haida Gwaii (; hai, X̱aaydag̱a Gwaay.yaay / , literally "Islands of the Haida people") is an archipelago located between off the northern Pacific coast of Canada. The islands are separated from the mainland to the east by the shallow Heca ...
(off western Canada)
* ''P. e. montana'' Ridgway, 1898 – inland southwestern Canada to west-central U.S.
* ''P. e. californica'' Price, 1897 – eastern California
* ''P. e. leucura'' ( Müller, PLS, 1776) – inland west, central Alaska to eastern Canada and northern New England (U.S.)
Description
This species is one of the largest species in the true finch
The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
family. It measures from in length and weighs from , with an average mass of . The pine grosbeak's wingspan is 13.0 in (33 cm). Among standard measurements, the wing chord is , the tail
The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...
is , the bill
Bill(s) may refer to:
Common meanings
* Banknote, paper cash (especially in the United States)
* Bill (law), a proposed law put before a legislature
* Invoice, commercial document issued by a seller to a buyer
* Bill, a bird or animal's beak
Plac ...
is and the tarsus is .[''CRC Handbook of Avian Body Masses'' by John B. Dunning Jr. (Editor). CRC Press (1992), .] Adults have a long forked black tail, black wings with white wing bars and a large bill. Adult males have a rose-red head, back and rump, They also possess black wings and tail, with a conical beak. Adult females are olive-yellow on the head and rump and grey on the back and underparts. Young birds have a less contrasting plumage
Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
overall, appearing shaggy when they moult
In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
their colored head plumage.
Its voice is geographically variable, and includes a whistled ''pui pui pui'' or ''chii-vli''. The song
A song is a musical composition intended to be performed by the human voice. This is often done at distinct and fixed pitches (melodies) using patterns of sound and silence. Songs contain various forms, such as those including the repetitio ...
is a short musical warble.
Distribution and habitat
Pine grosbeaks breed in the boreal forests of northern Eurasia and North America, and typically either remain resident near their breeding grounds or migrate relatively short distances to the southern extent of boreal forests. During irruptive years, more travel to southern boreal forests and some move further south. In such years in the New World, they can occur well south of their typical winter distribution, which is the northern Great Lakes region
The Great Lakes region of North America is a binational Canadian–American region that includes portions of the eight U.S. states of Illinois, Indiana, Michigan, Minnesota, New York, Ohio, Pennsylvania and Wisconsin along with the Canadian p ...
and northern New England
New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
in the United States. This species is a very rare vagrant to temperate
In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
parts of Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
; in all of Germany
Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
, for example, not more than 4 individuals per year and often none at all have been recorded since 1980. The birds have also been known to live in coniferous forests, and other woodlands of the sort.
Behaviour and ecology
The breeding habitat of the pine grosbeak is conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
ous forest
A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s. They nest on a horizontal branch or in a fork of a conifer. This bird is a permanent resident through most of its range; in the extreme north or when food sources are scarce, they may migrate farther south. When breeding both sexes develop which they use to store seeds before feeding them to the young.
Pine grosbeaks forage
Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also used m ...
in trees and bushes. They mainly eat seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s, buds, berries, and insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s. Outside of the nesting season, they often feed in flocks.
Pinicola enucleator5.jpg, Immature male
Pine Grosbeak, female.jpg, Front view of female, notice forked tail, Gatineau Park
Gatineau Park (french: Parc de la Gatineau) is a federal park located in the Outaouais, Outaouais region of Quebec, Canada. Administered by the National Capital Commission as part of the National Capital Region (Canada), National Capital Region, ...
, Quebec
Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirtee ...
Pinicola enucleator MHNT.ZOO.2010.11.220 Suomussalmi Naurois.jpg, Eggs of ''Pinicola enucleator'' MHNT
References
External links
*
Pine Grosbeak – ''Pinicola enucleator''
– USGS Patuxent Bird Identification InfoCenter
Xeno-canto: audio recordings of the pine grosbeak
*
{{Taxonbar, from=Q214571
pine grosbeak
The pine grosbeak (''Pinicola enucleator'') is a large member of the true finch family, Fringillidae. It is the only species in the genus ''Pinicola''. It is found in coniferous woods across Alaska, the western mountains of the United States, Can ...
pine grosbeak
The pine grosbeak (''Pinicola enucleator'') is a large member of the true finch family, Fringillidae. It is the only species in the genus ''Pinicola''. It is found in coniferous woods across Alaska, the western mountains of the United States, Can ...
Holarctic birds
pine grosbeak
The pine grosbeak (''Pinicola enucleator'') is a large member of the true finch family, Fringillidae. It is the only species in the genus ''Pinicola''. It is found in coniferous woods across Alaska, the western mountains of the United States, Can ...
pine grosbeak
The pine grosbeak (''Pinicola enucleator'') is a large member of the true finch family, Fringillidae. It is the only species in the genus ''Pinicola''. It is found in coniferous woods across Alaska, the western mountains of the United States, Can ...