botanical
Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
usage:
strobilus
A strobilus (plural: strobili) is a structure present on many land plant species consisting of sporangia-bearing structures densely aggregated along a stem. Strobili are often called cones, but some botanists restrict the use of the term cone to th ...
, plural strobili) is a seed-bearing organ on
gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνό ...
plants. It is usually woody, ovoid to globular, including scales and
bract
In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s arranged around a central axis, especially in
conifer
Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
s and
cycad
Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...
s. The cone of
Pinophyta
Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...
(conifer clade) contains the
reproductive
The reproductive system of an organism, also known as the genital system, is the biological system made up of all the anatomical organs involved in sexual reproduction. Many non-living substances such as fluids, hormones, and pheromones are als ...
structures. The woody cone is the female cone, which produces
seeds
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosperm pl ...
. The male cone, which produces
pollen
Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
, is usually
herbaceous
Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials.
Definitions of "herb" and "herbaceous"
The fourth edition of t ...
and much less conspicuous even at full maturity. The name "cone" derives from Greek ''konos'' (pine cone), which also gave name to the
geometric cone
A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex.
A cone is formed by a set of line segments, half-lines, or lines conn ...
. The individual plates of a cone are known as ''scales''. The ''umbo'' of a conifer cone refers to the first year's growth of a seed scale on the cone, showing up as a protuberance at the end of the two-year-old scale.
The male cone (microstrobilus or pollen cone) is structurally similar across all conifers, differing only in small ways (mostly in scale arrangement) from species to species. Extending out from a central axis are
microsporophyll
A sporophyll is a leaf that bears sporangia. Both microphylls and megaphylls can be sporophylls. In heterosporous plants, sporophylls (whether they are microphylls or megaphylls) bear either megasporangia and thus are called megasporophylls, or mi ...
s (modified leaves). Under each microsporophyll is one or several
microsporangia
Microsporangia are sporangia that produce microspores that give rise to male gametophytes when they germinate. Microsporangia occur in all vascular plants that have heterosporic life cycles, such as seed plants, spike mosses and the aquatic fer ...
(
pollen
Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
sacs).
The female cone (megastrobilus, seed cone, or ovulate cone) contains
ovule
In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...
s which, when fertilized by pollen, become seeds. The female cone structure varies more markedly between the different conifer families, and is often crucial for the identification of many species of conifers.
Female cone of the conifer families
Pinaceae cones
The members of the
pine family
The Pinaceae, or pine family, are conifer trees or shrubs, including many of the well-known conifers of commercial importance such as Cedrus, cedars, firs, Tsuga, hemlocks, larches, pines and spruces. The family is included in the order Pinales, ...
(
pine
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accep ...
s,
spruce
A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfami ...
s,
fir
Firs (''Abies'') are a genus of 48–56 species of evergreen coniferous trees in the family (biology), family Pinaceae. They are found on mountains throughout much of North America, North and Central America, Europe, Asia, and North Africa. The ...
s,
cedar
Cedar may refer to:
Trees and plants
*''Cedrus'', common English name cedar, an Old-World genus of coniferous trees in the plant family Pinaceae
*Cedar (plant), a list of trees and plants known as cedar
Places United States
* Cedar, Arizona
* ...
s,
larch
Larches are deciduous conifers in the genus ''Larix'', of the family Pinaceae (subfamily Laricoideae). Growing from tall, they are native to much of the cooler temperate northern hemisphere, on lowlands in the north and high on mountains furt ...
es, etc.) have cones that are imbricate (that is, with scales overlapping each other like fish scales). These pine cones, especially the woody female cones, are considered the "archetypal" tree cones.
The female cone has two types of scale: the
bract
In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
scales, and the seed scales (or ovuliferous scales), one
subtended
In geometry, an angle is subtended by an arc, line segment or any other section of a curve when its two rays pass through the endpoints of that arc, line segment or curve section. Conversely, the arc, line segment or curve section confined wi ...
by each bract scale, derived from a highly modified
branchlet
{{Short pages monitor
Image:Sapporo_Picea_Abies_Pinecone_1.jpg, Cluster of
Norway Spruce
''Picea abies'', the Norway spruce or European spruce, is a species of spruce native to Northern, Central and Eastern Europe.
It has branchlets that typically hang downwards, and the largest cones of any spruce, 9–17 cm long. It is very close ...
cones on
Hokkaidō
is Japan's second largest island and comprises the largest and northernmost prefecture, making up its own region. The Tsugaru Strait separates Hokkaidō from Honshu; the two islands are connected by the undersea railway Seikan Tunnel.
The la ...
.
Image:Arborvitae cone.png,
Arborvitae
''Thuja'' ( ) is a genus of coniferous tree or shrub in the Cupressaceae (cypress family). There are five species in the genus, two native to North America and three native to eastern Asia. The genus is monophyletic and sister to '' Thujopsis''. ...
cone.
Image:Pine Cones Jam.jpg, Pine cones jam (
Ukraine
Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inv ...
).
Image:Pinus canariensis (male) in Presa de las Niñas 02.jpg,
Pinus canariensis
''Pinus canariensis'', the Canary Island pine, is a species of gymnosperm in the conifer family Pinaceae. It is a large, evergreen tree, native and endemic to the outer Canary Islands of the Atlantic Ocean.
Description
''Pinus canariensis'' is a ...
male cone in
Gran Canaria
Gran Canaria (, ; ), also Grand Canary Island, is the third-largest and second-most-populous island of the Canary Islands, an archipelago off the Atlantic coast of Northwest Africa which is part of Spain. the island had a population of that co ...
.
File:Pine Cones.jpg, Pine Cones
File:More Pine Cones.jpg, More Pine Cones
File:Abies bracteata 01 Cone Peak.jpg, The cone structure of
Abies bracteata
''Abies bracteata'', the Santa Lucia fir or bristlecone fir, is the rarest and most endemic fir in North America, and according to some, the world. It is confined to steep-sided slopes and the bottoms of rocky canyons in the Santa Lucia Mountain ...
Visualisation of a ''Cupressus'' cone by micro computed tomography
File:Cupressus strobilus ("cone"), front and back.jpg, Photographs of the cone, front and back
File:ΜCT Cupressus strobilus 01.jpg, 3D visualisation of the cone
File:ΜCT Cupressus strobilus 02 - 3D section series.jpg, Various sections through the 3D object
File:ΜCT Cupressus strobilus 03 semi-transparent.jpg, Semi-transparent view
File:ΜCT Cupressus strobilus 04 mid section.jpg, Midsection
File:ΜCT of a cupressus strobilus, spiral 3D flight.ogg, Flight around the 3D object
File:ΜCT of a cupressus strobilus, frontal flight through image stack.ogg, Flight through the µCT image stack, lateral view
File:ΜCT of a cupressus strobilus, top flight through image stack.ogg, Flight through the µCT image stack, top view