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Armadillidiidae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
woodlice A woodlouse (plural woodlice) is an isopod crustacean from the polyphyleticThe current consensus is that Oniscidea is actually triphyletic suborder Oniscidea within the order Isopoda. They get their name from often being found in old wood. ...
, a
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
group in the order
Isopoda Isopoda is an order of crustaceans that includes woodlice and their relatives. Isopods live in the sea, in fresh water, or on land. All have rigid, segmented exoskeletons, two pairs of antennae, seven pairs of jointed limbs on the thorax, an ...
. Unlike members of some other woodlice families, members of this family can roll into a ball, an ability they share with the outwardly similar but unrelated
pill millipede Pill millipedes are any members of two living (and one extinct) orders of millipedes, often grouped together into a single superorder, Oniscomorpha. The name Oniscomorpha refers to the millipedes' resemblance to certain woodlice (Oniscidea), also ...
s and other animals. This ability gives woodlice in this family their
common names In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contra ...
of pill bugs or roly polies. Other common names include slaters, potato bugs, doodle bugs and cheeselogs. Most species are native to the Mediterranean Basin, while a few species have wider European distributions. The best-known species, ''
Armadillidium vulgare ''Armadillidium vulgare'', the common pill-bug, potato bug, common pill woodlouse, roly-poly, slater, doodle bug, or carpenter, is a widespread European species of woodlouse. It is the most extensively investigated terrestrial isopod species. ...
'', was introduced to
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
in the early 19th century and has become widespread throughout North America.


Ecology and behavior

Pill bugs in the family Armadillidiidae are able to form their bodies into a ball shape, in a process known as '' conglobation''. This behaviour is shared with
pill millipede Pill millipedes are any members of two living (and one extinct) orders of millipedes, often grouped together into a single superorder, Oniscomorpha. The name Oniscomorpha refers to the millipedes' resemblance to certain woodlice (Oniscidea), also ...
s (which are often confused with pill bugs),
armadillo Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along wi ...
s, and
cuckoo wasp Commonly known as cuckoo wasps or emerald wasps, the hymenopteran family Chrysididae is a very large cosmopolitan group (over 3000 described species) of parasitoid or kleptoparasitic wasps, often highly sculptured, with brilliant metallic colors ...
s. It may be triggered by
stimuli A stimulus is something that causes a physiological response. It may refer to: * Stimulation ** Stimulus (physiology), something external that influences an activity ** Stimulus (psychology), a concept in behaviorism and perception * Stimulus (eco ...
such as vibrations or pressure, and is a key defense against
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
; it may also reduce respiratory water losses. The diet of pill bugs is largely made up of decaying or decomposed plant matter such as leaves, and to a lesser extent, wood fibers. Pill bugs will also eat living plants, especially in wet conditions, sometimes consuming leaves, stems, shoots, roots, tubers, and fruits. Pill bugs can be serious pests in certain agricultural systems, particularly in areas that are prone to heavy rains and flood conditions. Pill bugs will feed on numerous crop plants including corn, beans, squash, peas, melon, chard, beet, cucumber, potato, spinach, lettuce, and strawberry, with potential for significant yield loss in strawberry in particular. Some species of pill bugs are known to eat decaying animal flesh or feces. They will also eat shed snakeskin and dead bugs, if necessary. Pill bugs contribute to their ecosystem as
decomposer Decomposers are organisms that break down dead or decaying organisms; they carry out decomposition, a process possible by only certain kingdoms, such as fungi. Like herbivores and predators, decomposers are heterotrophic, meaning that they use o ...
s. They are capable of taking in heavy metals such as copper, zinc, lead, and cadmium and crystallize these into spherical deposits in the
midgut The midgut is the portion of the embryo from which most of the intestines develop. After it bends around the superior mesenteric artery, it is called the "midgut loop". It comprises the portion of the alimentary canal from the end of the foregut ...
. They also provide a food source for birds, toads, spiders, wasps, and centipedes.


Classification

The family Armadillidiidae is differentiated from other woodlouse families by the two-segmented nature of the antennal flagellum, by the form of the
uropod Uropods are posterior appendages found on a wide variety of crustaceans. They typically have functions in locomotion. Definition Uropods are often defined as the appendages of the last body segment of a crustacean. An alternative definition sugge ...
s, and by the ability to roll into a ball. Within the family Armadillidiidae, 15 genera are currently recognized: *'' Alloschizidium'' Verhoeff, 1919 (13 species) *''
Armadillidium ''Armadillidium'' () is a genus of the small terrestrial crustacean known as the woodlouse. ''Armadillidium'' are also commonly known as pill woodlice, leg pebbles, pill bugs, roly-poly, or potato bugs, and are often confused with pill millipedes ...
'' Brandt, 1831 (189 species) *'' Ballodillium'' Vandel, 1961 (
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
) *'' Cristarmadillidium'' Arcangeli, 1936 (4 species) *'' Cyphodillidium'' Verhoeff, 1939 (monotypic) *'' Echinarmadillidium '' Verhoeff, 1901 (3 species) *'' Eleoniscus'' Racovitza, 1907 (monotypic) *'' Eluma'' Budde-Lund, 1885 (3 species) *'' Paraschizidium'' Verhoeff, 1919 (5 species) *'' Paxodillidium'' Schmalfuss, 1985 (monotypic) *'' Platanosphaera'' Strouhal, 1956 (6 species) *'' Schizidium'' Verhoeff, 1901 (26 species) *'' Trichodillidium'' Schmalfuss, 1989 (3 species) *'' Troglarmadillidium'' Verhoeff, 1900 (8 species) *'' Trogleluma'' Vandel, 1946 (2 species) *'' Typhlarmadillidium'' Verhoeff, 1900 (4 species) A 2022 study of
myrmecophilous Myrmecophily ( , ) is the term applied to positive interspecies associations between ants and a variety of other organisms, such as plants, other arthropods, and fungi. Myrmecophily refers to mutualistic associations with ants, though in its ...
populations indicated that these represented a new species of '' Cristarmadillidium'', and three new species within a new genus, '' Iberiarmadillidium''.


References


External links

* * * Regional maps for the most common American names for this isopod can be found in the results for question 74 o
the Harvard Dialect Survey
{{Taxonbar, from=Q1315837 Woodlice Crustacean families