Pileum
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''Pilea'', with 600–715 species, is the largest genus of flowering plants in the nettle family Urticaceae. It is distributed throughout the tropics, subtropics, and warm temperate regions (with the exception of
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
and New Zealand).


Description

The majority of species are
succulent In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meani ...
shade-loving herbaceous plants or
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s, which are easily distinguished from other Urticaceae by the combination of opposite leaves (with rare exceptions) with a single ligulate intrapetiolar stipule in each leaf axil and cymose or paniculate inflorescences (again with rare exceptions).


Uses

''Pilea'' is of little economic importance; one species is used in Chinese traditional medicine (''P. plataniflora'').


Horticulture

Six species have horticultural value (''P. cadierei'', ''P. grandifolia'', ''P. involucrata'', ''P. microphylla'', ''P. nummulariifolia'', and ''P. peperomioides''), Some pileas are grown for their ornamental foliage which is shaped like lily-pads. The ASPCA includes many peperomia species in the list of plants that are non-toxic to pets.


Systematics

The genus has attracted little monographic attention since Weddell (1869), and the majority of taxonomic contributions have come from floristic treatments. To date, 787 species names have been published (International Plant Names Index, 2003) and estimates for the species number range from 250 to 1000. Based on previous floristic treatments, about 30% of the species from regions not yet covered by contemporary floristic treatments may be undescribed. The genus name ''Pilea'' is Latin for "felt cap", a reference to the
calyx Calyx or calyce (plural "calyces"), from the Latin ''calix'' which itself comes from the Ancient Greek ''κάλυξ'' (''kálux'') meaning "husk" or "pod", may refer to: Biology * Calyx (anatomy), collective name for several cup-like structures ...
covering the
achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not ope ...
.


Selected species

*''
Pilea cadierei ''Pilea cadierei'', the aluminium plant or watermelon pilea, is a species of flowering plant in the nettle family Urticaceae, native to China (provinces of Guizhou and Yunnan) and Vietnam. The specific epithet ''cadierei'' refers to the 20th-cen ...
'' — aluminium plant *''
Pilea cataractae ''Pilea cataractae'' is a species of plant in the family Urticaceae. It is endemic to Mauritius. Its natural habitat is subtropical or tropical dry forest A forest is an area of land dominated by trees. Hundreds of definitions of forest ...
'' *''
Pilea cavernicola ''Pilea cavernicola'' is a herbaceous plant about 0.5 meters tall, native to China. A sciophyte, it grows in very low light conditions in caves in Fengshan County, Guangxi, China China, officially the People's Republic of China (PRC), is ...
'' *''
Pilea crassifolia ''Pilea crassifolia'' is a plant native to Central and South America. One well-known cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to ...
'' *''
Pilea depressa ''Pilea depressa'', the depressed clearweed, also sold as kiereweed, is a plant native to the Caribbean.Pilea de ...
'' *''
Pilea elegans ''Pilea elegans'' is a species in the plant family Urticaceae. It is found in Chile. USDA hardiness zone 9. The plant does not tolerate snow, but can tolerate occasional freezing spells of about -5 degrees C. Light requirements consist of indir ...
'' *''
Pilea fontana ''Pilea fontana'', the lesser clearweed, is an herbaceous plant which is very similar to ''Pilea pumila'' (both occupying an almost identical range covering most of North America east of the Rocky Mountains). They can be distinguished by the app ...
'' *''
Pilea glaucophylla ''Pilea glaucophylla'', the silver-leaved artillery plant, is a species of flowering plant in the family Urticaceae, native to Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regi ...
'' *''
Pilea grandifolia ''Pilea grandifolia'' is an undershrub native only to Jamaica. It is sometimes cultivated.
''National B ...
'' *''
Pilea involucrata ''Pilea involucrata'', commonly called the friendship plant, is a bushy trailing plant which is sometimes cultivated, especially where high humidity can be provided, such as in a terrarium A terrarium (plural: terraria or terrariums) is usua ...
'' — friendship plant *''
Pilea jamesonia ''Pilea jamesonia'' is a species of plant in the family Urticaceae. It is endemic to Ecuador. Its natural habitat is subtropical or tropical moist montane forests. References Endemic flora of Ecuador jamesonia ''Jamesonia'' is a g ...
'' *''
Pilea laevicaulis ''Pilea laevicaulis'' is a species of plant in the family Urticaceae. It is endemic to Mauritius. Its natural habitat is subtropical or tropical dry forest A forest is an area of land dominated by trees. Hundreds of definitions of forest ...
'' *'' Pilea microphylla'' — artillery plant, gunpowder plant *''
Pilea mollis ''Pilea mollis'' is a species of flowering plant in the family Urticaceae. It is used as an ornamental plant, particularly the cultivar 'Moon Valley'. References mollis Mollis is a former municipality in the canton of Glarus in Switzerla ...
'' — Moon Valley plant *'' Pilea myriophylla'' *''
Pilea napoana ''Pilea napoana'' is a species of plant in the family Urticaceae. It is endemic to Ecuador. Its natural habitat is subtropical or tropical moist montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in ...
'' *''
Pilea nummulariifolia ''Pilea nummulariifolia'' is a perennial evergreen herbaceous plant commonly known as creeping charlie native to the Caribbean (including Florida Florida is a state located in the Southeastern region of the United States. Florida is bord ...
'' — creeping Charlie *'' Pilea peperomioides'' — Chinese money plant, missionary plant *''
Pilea pollicaris ''Pilea pollicaris'' is a species of plant in the family Urticaceae. It is endemic to Mauritius. Its natural habitat is subtropical or tropical dry forest A forest is an area of land dominated by trees. Hundreds of definitions of forest ...
'' *''
Pilea pumila ''Pilea pumila'', commonly known as clearweed, Canadian clearweed, coolwort or richweed, is an herbaceous plant in the nettle family (Urticaceae). It is native to Asia and eastern North America, where it is broadly distributed. This plant is mos ...
'' — Canadian clearweed *''
Pilea repens ''Pilea repens'' is a plant which is sometimes cultivated, especially where high humidity can be provided, such as in a terrarium A terrarium (plural: terraria or terrariums) is usually a sealable glass container containing soil and plants th ...
'' — black-leaf panamiga *'' Pilea riopalenquensis'' *''
Pilea schimpfii ''Pilea schimpfii'' is a species of plant in the family Urticaceae. It is endemic to the Chimborazo, Napo, El Oro, and Carchi provinces of Ecuador. Its natural habitats are subtropical or tropical moist lowland forests and subtropical or tropica ...
'' *''
Pilea selbyanorum ''Pilea selbyanorum'' is a species of plant in the family Urticaceae. It is endemic to Ecuador. Its natural habitat is subtropical or tropical moist lowland forest A forest is an area of land dominated by trees. Hundreds of definitions ...
'' *''
Pilea serpyllacea ''Pilea serpyllacea'' is an herbaceous plant native to Venezuela, Bolivia, Colombia, Ecuador, and Peru. It has been introduced into a number of areas outside its native range, such as Hawaii Hawaii ( ; haw, Hawaii or ) is a state in ...
'' *''
Pilea serratifolia ''Pilea serratifolia'' is a species of plant in the family Urticaceae. It is endemic to Ecuador. Its natural habitats are subtropical or tropical moist montane forests and subtropical or tropical high-altitude shrubland Shrubland, scrubland ...
'' *''
Pilea spruceana ''Pilea spruceana'' (silver tree) is a species of evergreen plant in the family Urticaceae, which grows up to 1 foot in height, with a spread of up to 1.5 feet. Synonyms * ''Adicea spruceana'' (Wedd.) Kuntze References * ''Prodr.'' 16(1): 1 ...
'' *''
Pilea topensis ''Pilea topensis'' is a species of plant in the family Urticaceae. It is endemic to Ecuador. Its natural habitat is subtropical or tropical moist montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate in ...
'' *'' Pilea trianthemoides'' *''
Pilea trichosanthes ''Pilea trichosanthes'' is a species of plant in the family Urticaceae. It is endemic to Ecuador. The natural habitats of Philea trichosanthes are subtropical or tropical moist lowland forests and subtropical or tropical moist montane forests. ...
'' *''
Pilea trilobata ''Pilea trilobata'' is an endemic Mauritian plant from the genus ''Pilea'' within the family Urticaceae. It was first described by botanist Hugh Algernon Weddell in 1854. It was thought to be extinct since 1849 until it was rediscovered in April ...
'' *''
Pilea tungurahuae ''Pilea tungurahuae'' is a species of plant in the family Urticaceae. It is endemic to Ecuador. Its natural habitat is subtropical or tropical moist montane forest Montane ecosystems are found on the slopes of mountains. The alpine climate ...
'' *'' Pilea victoriae''


Fossil record

The fossil species †''Pilea cantalensis'' was widely distributed in Europe and
West Siberia Western Siberia or West Siberia (russian: Западная Сибирь, Zapadnaya Sibir'; kk, Батыс Сібір) is a part of the larger region of Siberia that is mostly located in the Russian Federation. It lies between the Ural region an ...
during the Miocene and Pliocene. It is related to the
East Asian East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. China, North Korea, South Korea a ...
''Pilea mongolica'' and to the North American ''
Pilea pumila ''Pilea pumila'', commonly known as clearweed, Canadian clearweed, coolwort or richweed, is an herbaceous plant in the nettle family (Urticaceae). It is native to Asia and eastern North America, where it is broadly distributed. This plant is mos ...
''.Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) zczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska) Acta Palaeobotanica 1979 20 (1): 3-117.


References

* * *Hortus Third, pages 872-873 * * * *
USDA U.S. Dept. of Agriculture Plants Database


Further reading

* * {{taxonbar , from=Q7222652 Urticaceae genera