The Mexican Plateau horned lizard (''Phrynosoma orbiculare'') is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

horned lizard Horned lizards (''Phrynosoma''), also known as horny toads or horntoads, are a genus of North American lizards and the type genus of the family Phrynosomatidae. The common names refer directly to their horns or to their flattened, rounded bodies, ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Phrynosomatidae The Phrynosomatidae are a diverse family of lizards, sometimes classified as a subfamily (Phrynosomatinae), found from Panama to the extreme south of Canada. Many members of the group are adapted to life in hot, sandy deserts, although the spiny ...

. The species, also known commonly as the Chihuahua Desert horned lizard, is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

. There are five recognized

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

. The specific epithet, ''orbiculare'', comes from the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

adjective ''orbis'', meaning "circular".

Description

''P. orbiculare'' has a characteristic single row of lateral abdominal fringe

scales Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number w ...

. This "horned toad" also has two short

occipital The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone overlies the occipital lobes of the cereb ...

horns Horns or The Horns may refer to: * Plural of Horn (instrument), a group of musical instruments all with a horn-shaped bells * The Horns (Colorado), a summit on Cheyenne Mountain * ''Horns'' (novel), a dark fantasy novel written in 2010 by Joe Hill ...

Geographic range

''P. orbiculare'' is found only in the high

plateau In geology and physical geography, a plateau (; ; ), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. Often one or more sides ha ...

country of central

. Specifically, it is found in the Mexican states of Chihuahua, Durango,

Morelos Morelos (), officially the Free and Sovereign State of Morelos ( es, Estado Libre y Soberano de Morelos), is one of the 32 states which comprise the Federal Entities of Mexico. It is divided into 36 municipalities and its capital city is Cuer ...

Nuevo León Nuevo León () is a state in the northeast region of Mexico. The state was named after the New Kingdom of León, an administrative territory from the Viceroyalty of New Spain, itself was named after the historic Spanish Kingdom of León. With a ...

Puebla Puebla ( en, colony, settlement), officially Free and Sovereign State of Puebla ( es, Estado Libre y Soberano de Puebla), is one of the 32 states which comprise the Federal Entities of Mexico. It is divided into 217 municipalities and its cap ...

, and

Veracruz Veracruz (), formally Veracruz de Ignacio de la Llave (), officially the Free and Sovereign State of Veracruz de Ignacio de la Llave ( es, Estado Libre y Soberano de Veracruz de Ignacio de la Llave), is one of the 31 states which, along with Me ...

Habitat

''P. orbiculare'' occurs in a wide range of primary

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s (dry scrubland, pine-oak forest, oak forest, juniper forest) and secondary habitats (agricultural land, and agave and '' Opuntia'' fields).

Subspecies

Five

of ''P. orbiculare'' are recognized as being valid, including the

nominotypical subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

. *''Phrynosoma orbiculare bradti'' *''Phrynosoma orbiculare cortezii'' *''Phrynosoma orbiculare dugesii'' *''Phrynosoma orbiculare orbiculare'' *''Phrynosoma orbiculare orientale'' ''

Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...

'': A trinomial authority in parentheses indicates that the subspecies was originally described in a genus other than ''Phrynosoma''.

Reproduction

''P. orbicularis'' is

viviparous Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the ...

Etymology

The

subspecific name In zoological nomenclature, a subspecific name is the third part of a trinomen. In zoology there is only one rank below that of species, namely "subspecies". In botanical nomenclature, there are several levels of subspecific names, such as ''var ...

, ''cortezii'', is in honor of Spanish ''conquistador''

Hernán Cortés Hernán Cortés de Monroy y Pizarro Altamirano, 1st Marquess of the Valley of Oaxaca (; ; 1485 – December 2, 1547) was a Spanish ''conquistador'' who led an expedition that caused the fall of the Aztec Empire and brought large portions of w ...

. The subspecific name, ''dugesii'', is in honor of French-born Mexican naturalist

Alfredo Dugès Alfredo Dugès (birth name- Alfred Auguste Delsescautz Dugès); (16 April 1826 – 7 January 1910) was a French-born, Mexican physician and naturalist born in Montpellier. He was the son of zoologist Antoine Louis Dugès (1797-1838). Alfredo Dugè ...

, who is considered the "father" of Mexican herpetology. Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Phrynosoma orbiculare cortezii'', p. 60; ''P. o. dugesii'', p. 76).