Phrynocephalus Helioscopus Horvathi
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''Phrynocephalus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
which includes 33
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of small and medium-sized agamid lizards, commonly called toadhead agamas or toad-headed agamas, that inhabit open arid and semiarid environments of
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
and
Eastern Europe Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russ ...
. The
systematics Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic tre ...
of this genus are very complicated with many controversial points of view about the unclear
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
of this group. All representatives of this genus have adopted the so-called "sit and wait" hunting strategy and they actively use visual orientation when watching for food. In general, the
ecological niche In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for ...
and role of ''Phrynocephalus'' species in lizard communities of arid environments of Asia are poorly studied, but seem to be similar to that of '' Phrynosoma,
Cophosaurus The greater earless lizard (''Cophosaurus texanus'') is the only species in the monotypic genus ''Cophosaurus''. It is closely related to the smaller, lesser earless lizards and other species in the genus ''Holbrookia'', and in fact was placed in ...
, Holbrookia,
Uta Uta or UTA may refer to: Universities *University of Texas at Arlington, in the United States *University of Tarapacá, in Chile *University of Tampere, in Finland Sports * FC UTA Arad, a Romanian football club based in the town of Arad * A c ...
'', and '' Sceloporus'' in the
New World The term ''New World'' is often used to mean the majority of Earth's Western Hemisphere, specifically the Americas."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: Oxford University Press, p. 3 ...
, as well as ''
Moloch Moloch (; ''Mōleḵ'' or הַמֹּלֶךְ‎ ''hamMōleḵ''; grc, Μόλοχ, la, Moloch; also Molech or Molek) is a name or a term which appears in the Hebrew Bible several times, primarily in the book of Leviticus. The Bible strongly co ...
'' in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
.


Species

The following 34
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are recognized as being valid.''Phrynocephalus''
The Reptile Database. www.reptile-database.org. *''
Phrynocephalus ananjevae Natalie's toad-headed agama (''Phrynocephalus ananjevae'') is a species of agamid lizard endemic to the Zagros Mountains in Iran. The specific epithet honors Natalia Ananjeva of the Zoological Institute in St. Petersburg, Russia for her contrib ...
'' – Natalie's toadhead agama *''
Phrynocephalus arabicus Arabian toad-headed agama (''Phrynocephalus arabicus'') is a species of agamid lizard found in Saudi Arabia, Qatar, United Arab Emirates, Oman, Iran, and Jordan. Description It is one of 34 recognized species within the genus ''Phrynocephalus' ...
'' – Arabian toadhead agama *''
Phrynocephalus axillaris Yarkand toad-headed agama (''Phrynocephalus axillaris''), is a species of agamid lizard found in Turkestan, Mongolia, and Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It ...
'' – Yarkand toadhead agama *''
Phrynocephalus clarkorum ''Phrynocephalus clarkorum'', also known commonly as the Afghan toad-headed agama and Clark's toad-headed agama, is a species of lizard in the family Agamidae. The species is native to parts of Central and South Asia. Etymology This species ...
'' – Afghan toadhead agama *''
Phrynocephalus erythrurus Sagus Kul lizard (''Phrynocephalus erythrurus'') is a species of agamid lizard endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; ...
'' – Sagus Kul lizard *''
Phrynocephalus euptilopus Alcock's toad-headed agama (''Phrynocephalus euptilopus''), is a species of agamid lizard found in Pakistan and Afghanistan. The species was originally established based on just 6 specimens from Darband (Kech), Darband, Western Baluchistan. Only ...
'' – Alcock's toadhead agama *''
Phrynocephalus forsythii ''Phrynocephalus forsythii'', also known commonly as Forsyth's toadhead agama and Forsyth's toad-headed lizard, is a species of lizard in the family Agamidae. The species is endemic to China. Etymology The specific name, ''forsythii'', is in h ...
'' –
Forsyth's R. W. Forsyth's often just called Forsyth's, is a group of Scottish department stores, the largest being at 26 - 30 Princes Street in Edinburgh. R. W. Forsyth Robert Wallace Forsyth was born Haddington, East Lothian in 1843 the son of Alexa ...
toadhead agama *''
Phrynocephalus frontalis ''Phrynocephalus frontalis'', the Shansi toadhead agama, is a species of agamid lizard endemic to China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a ...
'' –
Shansi Shanxi (; ; formerly romanised as Shansi) is a landlocked province of the People's Republic of China and is part of the North China region. The capital and largest city of the province is Taiyuan, while its next most populated prefecture-lev ...
toadhead agama *'' Phrynocephalus golubewii'' *'' Phrynocephalus guttatus'' – spotted toadhead agama *''
Phrynocephalus helioscopus ''Phrynocephalus helioscopus'', the sunwatcher toadhead agama, Fergana toad-headed agama, or sunwatcher, is a species of agamid lizard found in Kazakhstan, S Russia, Turkmenistan, Uzbekistan, Kyrgyzstan, Tajikistan, Turkey, Iraq, China, Mongolia, ...
'' – sunwatcher toadhead agama *'' Phrynocephalus hispidus'' – Dzhungar variegated toadhead agama *''
Phrynocephalus horvathi ''Phrynocephalus horvathi'' is a small diurnal desert lizard in the family Agamidae. It is endemic to the valley of the Aras River and considered Critically Endangered. Taxonomy The phylogeny within the genus ''Phrynocephalus'' is not well und ...
'' – Horvath's toadhead agama *'' Phrynocephalus interscapularis'' – Lichtenstein's toadhead agama *'' Phrynocephalus kulagini'' – Kulagin’s variegated toadhead agama *''
Phrynocephalus lutensis ''Phrynocephalus lutensis'', the Lut Desert toad-headed agama, is a species of agamid lizard found in Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bord ...
'' – Lut Desert toadhead agama *''
Phrynocephalus luteoguttatus ''Phrynocephalus luteoguttatus'', the yellow-speckled toad-headed agama, is a species of agamid lizard found in Iran, Afghanistan, Pakistan (Baluchistan) and possibly in India. The type locality is between Nushki and Helmand Helmand (Pashto/ ...
'' – yellow-speckled toadhead agama *''
Phrynocephalus maculatus ''Phrynocephalus maculatus'', the blacktail toadhead agama or spotted toad-headed agama, is a species of agamid lizard in Iran, Turkmenistan, Syria, Jordan, Iraq, Afghanistan, Pakistan, Saudi Arabia, Oman, United Arab Emirates, and Kuwait Ku ...
'' – blacktail toadhead agama *'' Phrynocephalus mystaceus'' – secret toadhead agama '' Phrynocephalus mystaceus'' in Astrakhan Oblast, Russia.">Russia.html" ;"title="Astrakhan Oblast, Russia">Astrakhan Oblast, Russia. *''Phrynocephalus ornatus'' *''Phrynocephalus persicus'' – Persian toadhead agama *''Phrynocephalus przewalskii'' – Nikolay Przhevalsky, Przewalski's toadhead agama *''Phrynocephalus putjatai'' *''Phrynocephalus raddei'' *''
Phrynocephalus reticulatus The reticulated toad-headed agama (''Phrynocephalus reticulatus'') is a species of agamid lizard found in Central Asia and West Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of ...
'' – reticulated toadhead agama *'' Phrynocephalus roborowskii'' – Roborowski's toadhead agama *'' Phrynocephalus rossikowi'' –
Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked cou ...
toadhead agama *''
Phrynocephalus sakoi ''Phrynocephalus sakoi'' is a species of agamid lizard endemic to Oman Oman ( ; ar, عُمَان ' ), officially the Sultanate of Oman ( ar, سلْطنةُ عُمان ), is an Arabian country located in southwestern Asia. It is situated on ...
'' *''
Phrynocephalus scutellatus ''Phrynocephalus scutellatus'', the gray toadhead agama or gray toad head agama, is a species of agamid lizard. It is found in Iran, Afghanistan, and southwestern Pakistan. References

Phrynocephalus, scutellatus Reptiles of Afghanistan ...
'' – gray toadhead agama *'' Phrynocephalus strauchi'' – Strauch's toadhead agama *''
Phrynocephalus theobaldi ''Phrynocephalus theobaldi'' (common names: Theobald's toad-headed agama, snow lizard, and others) is a species of lizard in the Family (biology), family Agamidae. The species is Endemism, endemic to Asia. Etymology The Specific name (zoology), ...
'' – Theobald's toadhead agama, toad-headed lizard, or snow lizard *''
Phrynocephalus versicolor ''Phrynocephalus versicolor'', the Tuvan toad-headed agama, is a species of agamid lizard found in desert areas of China and Mongolia. It was first described by the Russian herpetologist Alexander Strauch, director of the Zoological Museum at th ...
'' – Tuvan toadhead agama *'' Phrynocephalus vindumi'' *'' Phrynocephalus vlangalii'' –
Ching Hai Ching Hai (born Trịnh Đăng Huệ;Vietnamese name consisting of three parts in the following order: a family name, a middle name and a given name. 12 May 1950), commonly referred to as ''Suma'' or '' Supreme Master Ching Hai (Chinese: 清海 ...
toadhead agama ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A
binomial authority In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
in parentheses indicates that the species was originally described in a genus other than ''Phrynocephalus''.


References


Further reading

" Caup Kaup JJ (1825). "''Einige Bemerkungen zu Merrems Handbuch'' Merrem's Handbook">Blasius_Merrem.html" ;"title=" Some Remarks on Blasius Merrem">Merrem's Handbook. ''Isis von Oken'' 16: 589-593. (''Phrynocephalus'', new genus, p. 591). (in German). {{Taxonbar">from=Q1790689 Lizard genera Taxa named by Johann Jakob Kaup