Pholad borings are tubular burrows in firm clay and soft rock that have been created by

bivalve Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bival ...

molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant taxon, extant species of molluscs are recognized. The number of fossil sp ...

(boring clams) in the family

Pholadidae Pholadidae, known as piddocks or angelwings, are a family of bivalve molluscs similar to a clam. Background Piddocks are unique in that each side of their shells is divided into 2 or 3 separate sections. Furthermore, one of the piddock's shells ...

. The

common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...

s of clams in this family are "pholads", "piddocks", and "angel wings"; the latter because their shells are white, elongated and tend to be shaped like a wing and have sculpture somewhat reminiscent of a wing. Pholads are unusual bivalves capable of boring into various kinds of rock, firm clay and peat, and they live permanently in the burrows they create. These clams use a set of ridges or "teeth" on the outer anterior surfaces of their shells to grind into suitable substrate, creating a non-branching burrow. Pholad burrows are typically narrower at the entrance of the burrow than they are across the interior. Ancient, sand-filled pholad borings are a distinctive type of

trace fossil A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, ...

known as '' Gastrochaenolites'' that are found in some

limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...

dolomite Dolomite may refer to: *Dolomite (mineral), a carbonate mineral *Dolomite (rock), also known as dolostone, a sedimentary carbonate rock *Dolomite, Alabama, United States, an unincorporated community *Dolomite, California, United States, an unincor ...

s and lithified

mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from '' shale'' by its lack of fissility (parallel layering).Blatt, H., and R.J. Tracy, 1996, ''Petrology. ...

s. They are assigned to the

Trypanites ''Trypanites'' is a narrow, cylindrical, unbranched boring which is one of the most common trace fossils in hard substrates such as rocks, carbonate hardgrounds and shells (Bromley, 1972). It appears first in the Lower Cambrian (James et al ...

ichnofacies An ichnofacies is an assemblage of trace fossils that provides an indication of the conditions that their formative organisms inhabited. Concept Trace fossil assemblages are far from random; the range of fossils recorded in association is constra ...

, and evidence submarine

hardgrounds Carbonate hardgrounds are surfaces of synsedimentarily cemented carbonate layers that have been exposed on the seafloor (Wilson and Palmer, 1992). A hardground is essentially, then, a lithified seafloor. Ancient hardgrounds are found in limestone ...

that formed soon after deposition, typically during episodes of non-sedimentation, with subsequent infilling of the burrow and burial when

sedimentation Sedimentation is the deposition of sediments. It takes place when particles in suspension settle out of the fluid in which they are entrained and come to rest against a barrier. This is due to their motion through the fluid in response to the ...

resumed. Pholad burrow trace fossils are not to be confused with specimens of the ''

ichnogenus An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...

, which are toothpick-sized masses of holes that characterize some hardgrounds. ''Trypanites'' probably result from a combination of mechanical abrasion and acid dissolution by the organism, whereas pholad borings are thought to be purely mechanical in origin.

References

Boring fossils Invertebrate paleozoology {{trace-fossil-stub

See also

References