Philip Hunter Timberlake
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Philip Hunter Timberlake (1883-1981) was one of the most prolific American entomologists of the 20th century. He was born on June 5, 1883 in
Bethel, Maine Bethel is a town in Oxford County, Maine, United States. The population was 2,504 at the 2020 census. It includes the villages of Bethel and West Bethel. The town is home to Gould Academy, a private preparatory school, and is near the Sun ...
, and died in 1981 in
Riverside, California Riverside is a city in and the county seat of Riverside County, California, United States, in the Inland Empire metropolitan area. It is named for its location beside the Santa Ana River. It is the most populous city in the Inland Empire an ...
, where he had served as an Associate Entomologist in the Department of Entomology of the
University of California, Riverside The University of California, Riverside (UCR or UC Riverside) is a public land-grant research university in Riverside, California. It is one of the ten campuses of the University of California system. The main campus sits on in a suburban distr ...
. He obtained an A.B. degree in 1908 in Liberal Arts from Bowdoin College with a major in Greek and Latin. In 1910 he received an A.M. degree in Biology from
Harvard University Harvard University is a private Ivy League research university in Cambridge, Massachusetts. Founded in 1636 as Harvard College and named for its first benefactor, the Puritan clergyman John Harvard, it is the oldest institution of high ...
. From 1909-1914, Timberlake was employed by the
U.S. Department of Agriculture The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of comme ...
Bureau of Entomology, as “Agent and Expert” conducting research in
biological control Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also i ...
of pest insects. From 1914-1924 he was Associate Entomologist at the Hawaiian Sugar Planters Experiment Station in
Honolulu Honolulu (; ) is the capital and largest city of the U.S. state of Hawaii, which is in the Pacific Ocean. It is an unincorporated county seat of the consolidated City and County of Honolulu, situated along the southeast coast of the island ...
, where his research dealt primarily with biological control using
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s and
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s. In 1924 he was appointed Associate Entomologist in the Department of Biological Control at the
Citrus Experiment Station The University of California Citrus Experiment Station is the founding unit of the University of California, Riverside campus in Riverside, California, United States. The station contributed greatly to the cultivation of the orange and the overall ...
of the University of California, Riverside, where he served until retirement in 1950. He continued his regular work schedule afterward, but his eyesight slowly failed in later years, and in 1980, at age 97 years, it reached the point where he could no longer continue his work. The appointment of Timberlake in the Department of Biological Control in 1924 was motivated by his extensive knowledge of the taxonomy of parasitic Hymenoptera and of predaceous ladybird beetles, groups of importance in biological control of pest insects. However, by the late 1920s and thereafter, he focused almost entirely upon the
taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
of native bees, especially the genus '' Perdita''. There are over 800 species of ''Perdita'', most described by Timberlake, but also by T.D.A. Cockerell. They are almost all specialist pollinators (
oligolege The term oligolecty is used in pollination ecology to refer to bees that exhibit a narrow, specialized preference for pollen sources, typically to a single family or genus of flowering plants. The preference may occasionally extend broadly to mu ...
s) of many species of plants, especially in the
Sonoran desert The Sonoran Desert ( es, Desierto de Sonora) is a desert in North America and ecoregion that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the southwestern United States (in Ariz ...
, where Timberlake carried out extensive collecting for decades. He described and named about 800 species of bees in total over his career, and several other species in other insect groups. He published over 100 scientific papers, mostly on bees, in addition to 8 volumes on the genus ''Perdita'' alone. His colleagues and former students throughout the world have described over 50 new species of insects, named ''timberlakei'' as patronyms in his honor. His insect collection contained about 500,000 specimens of which about 150,000 were Hymenoptera, including what was once the largest bee collection in North America, and this served as the foundation for the collection now housed in the University of California's
Entomology Research Museum The UCR Entomology Research Museum is the insect collection of the Department of Entomology of the University of California, Riverside. It contains approximately 4 million total insect specimens, over 3 million of which are pinned, roughly 400,0 ...
, containing some 4 million total specimens. He married his wife, Edith Timberlake, in 1917 in
Honolulu Honolulu (; ) is the capital and largest city of the U.S. state of Hawaii, which is in the Pacific Ocean. It is an unincorporated county seat of the consolidated City and County of Honolulu, situated along the southeast coast of the island ...
, and they had three children. Edith was the sister of Hannah Milhous Nixon and an aunt of former President Richard M. Nixon, whose law school career she helped finance. She and the elderly scientist attended Nixon's Presidential inauguration at his invitation, and she died a few years later, in 1972

{{DEFAULTSORT:Timerlake, Philip Hunter 1883 births 1981 deaths American entomologists Hymenopterists Bowdoin College alumni Harvard University alumni University of California, Riverside faculty 20th-century American zoologists