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''Phasianella'',
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
the pheasant shells, is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of small
sea snail Sea snail is a common name for slow-moving marine gastropod molluscs, usually with visible external shells, such as whelk or abalone. They share the taxonomic class Gastropoda with slugs, which are distinguished from snails primarily by the ...
s with a
calcareous Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adje ...
operculum and a colorfully patterned shell, a marine
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
in the family
Phasianellidae Phasianellidae common name the "pheasant shells" or "pheasant snails" is a family of small sea snails with calcareous opercula, marine gastropod mollusks in the subclass Vetigastropoda Vetigastropoda is a major Taxonomy (biology), taxonomi ...
.


Description

The white shell has a bulimiform or subglobose shape. It is polished, without epidermis or nacre, variegated with bright colors. The heavy operculum is calcareous, internally paucispiral, with a nucleus near the basal margin. It is externally convex. The
columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved and forms an important source on Roman agriculture, together with the wo ...
is smooth, arcuate and not dentate. The animal has long tentacles, and has usually pectinated head lobes. These frontal lobes distinguish this genus from '' Tricolia'', which lacks frontal lobes. The epipodial line contains generally cirrhi. The branchial plume is long. The narrow foot is long, pointed posteriorly, rounded before. Below it is divided longitudinally by a median groove. The rhomboidal jaws are covered with imbricating scales. The
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
is rather short. The formula of teeth is typically ∞ 5.1.5 ∞, but sometimes lacks the median and outer lateral teeth. Contrary to most spiral species, the two gills in Phasiannela are close together, so as to appear almost as one.


Distribution

The species in this genus occur in the tropical
Indo-West Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
and off
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
.


Species

Species within the genus ''Phasianella'' include: * ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' (Gmelin, 1791) * † '' Phasianella brandenburgi'' O. Boettger, 1907Robertson R. (2010). "A true ''Phasianella'' from the Middle Miocene (Badenian) Central Paratethys of Romania (Gastropoda: Vetigastropoda: Phasianellidae)". ''
Archiv für Molluskenkunde ''Archiv für Molluskenkunde'' is a peer-reviewed scientific journal published by the Senckenberg Nature Research Society, covering research in malacology. Coverage ''Archiv für Molluskenkunde'' publishes original research on all aspects of mo ...
'' 139(2): 247-253. .
* '' Phasianella bryani'' Pilsbry, 1917Henry A. Pilsbry (1917), Marine Mollusks of Hawaii, I-III; Proceedings of the Academy of Natural Sciences of Philadelphia, Vol. 69, No. 2 (Apr. -Sep., 1917), pp. 207-230
/ref> * '' Phasianella molokaiensis'' Pilsbry, 1917 (taxon inquirendum) * '' Phasianella nivosa'' Reeve, 1862 * '' Phasianella solida'' (Born, 1778) * '' Phasianella ventricosa'' Swainson, 1822 * '' Phasianella wisemanni'' Baird, 1873 (taxon inquirendum) ;Species brought into synonymy: * ''Phasianella adamsi'' Philippi, 1853: synonym of '' Eulithidium adamsi'' (Philippi, 1853) * ''Phasianella aethiopica'' Philippi, 1853: synonym of '' Phasianella solida'' (Born, 1778) * ''Phasianella affinis'' C. B. Adams, 1850: synonym of '' Eulithidium affine'' (C. B. Adams, 1850) * ''Phasianella africana'' Bartsch, 1915: synonym of '' Tricolia africana'' (Bartsch, 1915) * ''Phasianella angulifera'' Lamarck, 1822: synonym of '' Littoraria angulifera'' (Lamarck, 1822) * ''Phasianella bella'' M. Smith, 1937: synonym of '' Eulithidium bellum'' (M. Smith, 1937) * ''Phasianella bifasciata'' Brown, 1827: synonym of '' Lacuna vincta'' (Montagu, 1803) * ''Phasianella brevis'' C. B. Adams, 1850: synonym of '' Eulithidium adamsi'' (Philippi, 1853) * ''Phasianella brevis'' Vélain, 1877: synonym of '' Tricolia munieri'' (Vélain, 1877) * ''Phasianella brevis'' auct. non d'Orbigny, 1842: synonym of '' Gabrielona sulcifera'' Robertson, 1973 * ''Phasianella brevis'' Menke, 1843 : synonym of ''Phasianella ventricosa'' Swainson, 1822 (invalid: junior homonym of ''Phasianella brevis'' d'Orbigny, 1842; ''P. turgida'' is a replacement name) * ''Phasianella brevissima'' Pilsbry, 1888: synonym of '' Eulithidium breve'' (d'Orbigny, 1842) omen dubium* ''Phasianella bulimoides'' Lamarck, 1822: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' Gmelin, 1788 * ''Phasianella caloundra Iredale, 1927:OBIS: ''Phasianella caloundra''
/ref> synonym of '' Phasianella solida'' (Born, 1778) * ''Phasianella canalis'' Montagu, 1803: synonym of '' Lacuna vincta'' (Montagu, 1803) * ''Phasianella capensis'' Dunker, 1846: synonym of '' Tricolia capensis'' (Dunker, 1846) * ''Phasianella concinna'' C. B. Adams, 1850 : synonym of '' Eulithidium affine'' (C. B. Adams, 1850) * ''Phasianella concolor'' C. B. Adams, 1850: synonym of '' Assiminea succinea'' (Pfeiffer, 1840) * ''Phasianella cornea'' Brown, 1827: synonym of '' Lacuna vincta'' (Montagu, 1803) * ''Phasianella crassior'' Montagu, 1803: synonym of '' Lacuna crassior'' (Montagu, 1803) * ''Phasianella decorata'' Chenu: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' Gmelin, 1788 * ''Phasianella delicatula'' Tenison-Woods, J.E., 1877: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' Gmelin, 1788 * ''Phasianella elachista'' Melvill, 1901: synonym of '' Tricolia fordiana'' (Pilsbry, 1888) * ''Phasianella elongata'' Krauss, 1848: : synonym of '' Tricolia elongata'' (Krauss, 1848) * ''Phasianella fasciata'' Menke, 1830: synonym of ''
Bankivia fasciata ''Bankivia fasciata'', common name the banded kelp shell, is a species of sea snail, a marine gastropod mollusk in the family Trochidae, the top snails.Bouchet, P. (2012). ''Bankivia fasciata'' (Menke, 1830). Accessed through: World Register o ...
'' (Menke, 1830) * ''Phasianella ferussaci'' Guerin 1829 : synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella flammulata'' (Hutton, 1878): synonym of '' Eatoniella huttoni'' (Pilsbry, 1888) * ''Phasianella formosa'' Turton, 1932: synonym of '' Tricolia formosa'' (Turton, 1932) * ''Phasianella fulgurata'' Reeve, 1862: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella fulminata'' Menke, 1830: synonym of ''
Bankivia fasciata ''Bankivia fasciata'', common name the banded kelp shell, is a species of sea snail, a marine gastropod mollusk in the family Trochidae, the top snails.Bouchet, P. (2012). ''Bankivia fasciata'' (Menke, 1830). Accessed through: World Register o ...
'' (Menke, 1830) * ''Phasianella grata'' Philippi, 1853: synonym of '' Phasianella solida'' (Born, 1778) * ''Phasianella histrio'' Reeve, 1862: synonym of '' Phasianella solida'' (Born, 1778) * ''Phasianella inflexa'' Blainville, 1825: synonym of ''
Melanella dufresnii ''Melanella'' is a genus of very small ectoparasitic sea snails, marine gastropod mollusks or micromollusks in the family Eulimidae.Gofas, S. (2011). Melanella Bowdich, 1822. Accessed through: World Register of Marine Species at http://ww ...
'' Bowdich, 1822 * ''Phasianella intermedia'' Scacchi, 1836: synonym of '' Tricolia tenuis'' (Michaud, 1829) * ''Phasianella jaspidea'' Reeve, 1862: synonym of '' Phasianella solida'' (Born, 1778) * ''Phasianella kraussi'' E. A. Smith, 1911: synonym of '' Tricolia kraussi'' (E. A. Smith, 1911) * ''Phasianella lehmanni'' Menke, 1830: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' Gmelin, 1788 * ''Phasianella lentiginosa'' Reeve: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella lineata'' Lamarck, 1822: synonym of '' Littoraria tessellata'' (Philippi, 1847) * ''Phasianella lineata'' auct. non Lamarck, 1822: synonym of ''
Echinolittorina interrupta ''Echinolittorina interrupta'' is a species of sea snail, a marine gastropod mollusk in the family Littorinidae, the winkles or periwinkles. Distribution Description The maximum recorded shell Shell may refer to: Architecture and design ...
'' (Philippi, 1847) * ''Phasianella lineolata'' Wood, 1828: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella lipeata'' Krebs, 1864: synonym of '' Eulithidium tessellatum'' (Potiez & Michaud, 1838) * ''Phasianella lymnaeoides'' Anton 1839: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella meleagris'' Potiez & Michaud, 1838: synonym of '' Echinolittorina meleagris'' (Potiez & Michaud, 1838) * ''Phasianella minima'' Melvill, 1896: synonym of '' Tricolia fordiana'' (Pilsbry, 1888) * ''Phasianella minuta'' Anton, 1838: synonym of '' Eulithidium tessellatum'' (Potiez & Michaud, 1838) * ''Phasianella modesta'' Gould, 1861: synonym of '' Phasianella solida'' (Born, 1778) * ''Phasianella montebelloensis'' Preston, 1914: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella munieri'' Vélain, 1877: synonym of '' Tricolia munieri'' (Vélain, 1877) * ''Phasianella nebulosa'' Lamarck, 1822: synonym of '' Littoraria nebulosa'' (Lamarck, 1822) * ''Phasianella nepeanensis'' Gatliff & Gabriel, 1908: synonym of '' Gabrielona nepeanensis'' (Gatliff & Gabriel, 1908) * ''Phasianella neritina'' Dunker, 1846: synonym of '' Tricolia neritina'' (Dunker, 1846) * ''Phasianella nivosa'' Reeve, 1848: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella peruviana'' Lamarck, 1822: synonym of '' Echinolittorina peruviana'' (Lamarck, 1822) * ''Phasianella petiti'' Craven, 1882: synonym of '' Tricolia petiti'' (Craven, 1882) * ''Phasianella picta'' DeBlain: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' Gmelin, 1788 * ''Phasianella preissi'' Menke, 1830: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' Gmelin, 1788 * ''Phasianella pullus'' (Linnaeus, 1758): synonym of '' Tricolia pullus'' (Linnaeus, 1758) * ''Phasianella punctata'' Pfeiffer, 1840: synonym of '' Echinolittorina meleagris'' (Potiez & Michaud, 1838) * ''Phasianella reticulata'' Reeve, 1862: synonym of '' Phasianella ventricosa'' Swainson, 1822 * ''Phasianella rubens'' Lamarck, 1822: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella sanguinea'' Reeve, 1862: synonym of '' Phasianella ventricosa'' Swainson, 1822 * ''Phasianella speciosa'' var. ''albina'' Monterosato 1880: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''atra'' Bucquoy, Dautzenberg & Dollfus 1884: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''atrata'' Dautzenberg 1883: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''aurea'' Dautzenberg 1882: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''lactea'' Monterosato 1880: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''maculata'' Monterosato 1880: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''major'' Monterosato 1880: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''normalis'' Monterosato 1880: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''purpurea'' Dautzenberg 1883: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''sanguinea'' Bucquoy, Dautzenberg & Dollfus 1884: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''spirolineata'' Monterosato 1880: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella speciosa'' var. ''viridis'' Monterosato 1880: synonym of '' Tricolia speciosa'' (Mühlfeld, 1824) * ''Phasianella splendida'' Philippi, 1853: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella striata'' Brown, 1827: synonym of '' Lacuna vincta'' (Montagu, 1803) * ''Phasianella striolata'' Turton, 1932: synonym of '' Tricolia striolata'' (Turton, 1932) * ''Phasianella stylifera'' Turton, 1825: synonym of '' Pelseneeria stylifera'' (Turton, 1825) * ''Phasianella sulcata'' Lamarck, 1822: synonym of ''
Littoraria irrorata ''Littoraria irrorata'', also known by the common name the marsh periwinkle, is a species of sea snail, a marine gastropod mollusk in the family Littorinidae. The specific epithet ' means 'moistened' or 'dewy.' This species occurs in salt marshe ...
'' (Say, 1822) * ''Phasianella sulcosa'' Mighels, 1843: synonym of '' Odostomia sulcosa'' (Mighels, 1843) * ''Phasianella tessellata'' Potiez & Michaud, 1838: synonym of '' Eulithidium tessellatum'' (Potiez & Michaud, 1838) * ''Phasianella tesselata'' C. B. Adams, 1850: synonym of '' Eulithidium tessellatum'' (Potiez & Michaud, 1838) * ''Phasianella tessellata'' Anton, 1838: synonym of '' Echinolittorina punctata'' (Gmelin, 1791) * ''Phasianella thaanumi'' Pilsbry, 1917: synonym of '' Hiloa variabilis'' (Pease, 1861) * ''Phasianella tomlini'' Gatliff & Gabriel, 1921: synonym of '' Tricolia tomlini'' (Gatliff & Gabriel, 1921) * ''Phasianella tritonis'' Chemnitz, 1788: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' (Gmelin, 1791) * ''Phasianella turgida'' Philippi, 1853 : synonym of '' Phasianella ventricosa'' Swainson, 1822 * ''Phasianella undatella'' Menke, 1830: synonym of ''
Bankivia fasciata ''Bankivia fasciata'', common name the banded kelp shell, is a species of sea snail, a marine gastropod mollusk in the family Trochidae, the top snails.Bouchet, P. (2012). ''Bankivia fasciata'' (Menke, 1830). Accessed through: World Register o ...
'' (Menke, 1830) * ''Phasianella variabilis'' Pease, 1888: synonym of '' Hiloa variabilis'' (Pease, 1861) (unaccepted combination) * ''Phasianella variegata'' Lamarck, 1822: synonym of '' Phasianella solida'' (Born, 1778) (invalid: junior homonym of ''P. variegata'' de Roissy, 1805) * ''Phasianella velaini'' Lamy, 1931: synonym of '' Tricolia munieri'' (Vélain, 1877) * ''Phasianella umbilicatum'' auct. non d'Orbigny, 1842: synonym of '' Eulithidium affine'' (C. B. Adams, 1850) * ''Phasianella varia'' Lamarck, 1822: : synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' Gmelin, 1788 * ''Phasianella venosa'' Reeve, 1862: synonym of '' Phasianella ventricosa''Swainson, 1822 * ''Phasianella venusta'' Reeve, 1848: synonym of ''
Phasianella australis ''Phasianella australis'', common names the Australian pheasant, painted lady, and pheasant snail, is a medium-sized to large species of sea snail with a calcareous operculum and a colorfully patterned shell, a marine gastropod mollusc in t ...
'' (Gmelin, 1791) * ''Phasianella vincta'' Montagu, 1803: synonym of '' Lacuna vincta'' (Montagu, 1803) * ''Phasianella viridis'' Anton: synonym of '' Phasianella variegata'' Lamarck, 1822 * ''Phasianella zebra'' Gray in Reeve, 1862: synonym of '' Phasianella ventricosa''Swainson, 1822 * ''Phasianella zigzag'' Odhner, 1919: : synonym of '' Phasianella solida'' (Born, 1778)


References

* Vaught, K.C. (1989). ''A classification of the living Mollusca''. American Malacologists: Melbourne, FL (USA). . XII, 195 pp.


External links


Uniprot taxonomy

WoRMS
{{Commons category Phasianellidae Extant Miocene first appearances Gastropod genera Taxa named by Jean-Baptiste Lamarck