Petrophile Pulchella
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''Petrophile pulchella'', commonly known as conesticks, is a common shrub of the family Proteaceae and is found in eastern Australia. The leaves are divided with needle-shaped but soft
pinnae The auricle or auricula is the visible part of the ear that is outside the head. It is also called the pinna (Latin for "wing" or " fin", plural pinnae), a term that is used more in zoology. Structure The diagram shows the shape and location ...
, the flowers silky-hairy, cream-coloured and arranged in oval heads and the fruit are arranged in oval heads. Conesticks grows on shallow sandstone soils, often in open forest or
heathland A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a coole ...
s near the coast. It is also occasionally seen on the adjacent ranges.


Description

''Petrophile pulchella'' is a shrub that typically grows to a height of in sheltered locations but to only in exposed heathland. The branchlets and leaves are softly-hairy at first but become
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
with age. The leaves are long on a petiole long, and divided two or three times with needle-shaped pinnae but that are soft rather than sharp-tipped. The flowers are arranged in leaf axils and on the ends of branchlets in oval heads long and are
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
or on a peduncle up to about long. The flowers are long, cream-coloured and silky-hairy. Flowering mostly occurs from August to March and the fruit is a
nut Nut often refers to: * Nut (fruit), fruit composed of a hard shell and a seed, or a collective noun for dry and edible fruits or seeds * Nut (hardware), fastener used with a bolt Nut or Nuts may also refer to: Arts, entertainment, and media Co ...
long, fused with others in an oval head up to long. This species is distinguished from '' P. pedunculata'' by its flowerheads that are on peduncles long. The two other species in eastern Australia, '' P. canescens'' and '' P. sessilis'', both have finely hairy new growth.


Taxonomy

The shrub was first formally described in 1796 by Heinrich Schrader and
Johann Christoph Wendland Johann Christoph Wendland (July 17, 1755 – July 27, 1828) was a German botanist and gardener born in Petit-Landau, Alsace. Family His son, Heinrich Ludolph Wendland (1791–1869), and his grandson, Hermann Wendland (1825–1903), ...
who gave it the name ''Protea pulchella'' in ''Sertum Hannoveranum'', from the original specimen collected at
Botany Bay Botany Bay (Dharawal: ''Kamay''), an open oceanic embayment, is located in Sydney, New South Wales, Australia, south of the Sydney central business district. Its source is the confluence of the Georges River at Taren Point and the Cook ...
. In 1810, the prolific botanist Robert Brown reclassified it in the new genus ''Petrophile'' as ''P. pulchella'', publishing the name change in ''
Transactions of the Linnean Society of London The Linnean Society of London is a learned society dedicated to the study and dissemination of information concerning natural history, evolution, and taxonomy. It possesses several important biological specimen, manuscript and literature col ...
''. The specific epithet ''pulchella'' meaning “beautiful” is derived from Latin, although noted plant author John Wrigley feels it to be somewhat of a misnomer. Joseph Knight, who had propagated and cultivated it successfully in England by 1809, reported, "It has few claims to a place in our collections."


Distribution and habitat

''Petrophile pulchella'' is found from south-eastern Queensland and south along the coast and adjacent tablelands to
Jervis Bay Jervis Bay () is a oceanic bay and village on the south coast of New South Wales, Australia, said to possess the whitest sand in the world. A area of land around the southern headland of the bay is a territory of the Commonwealth of Australia ...
in New South Wales. It often grows with trees such as Sydney peppermint (''
Eucalyptus piperita ''Eucalyptus piperita'', commonly known as Sydney peppermint and urn-fruited peppermint, is a small to medium forest tree native to New South Wales, Australia. Description It has grey, rough and finely fibrous bark on its trunk, but its branche ...
''), smooth-barked apple ('' Angophora costata'') or more open woodland e.g. with scribbly gum (''
Eucalyptus sclerophylla ''Eucalyptus sclerophylla'', known as the scribbly gum, is a tree native to eastern Australia. Very similar to the related Scribbly Gum (''E. haemastoma''), a better known tree. The best way of distinguishing the species is the smaller hemisph ...
''), silvertop ash ('' E. sieberi'') or with shrubs such as mountain devil ('' Lambertia formosa''), broad-leaved drumsticks (''
Isopogon anemonifolius ''Isopogon anemonifolius'', commonly known as broad-leaved drumsticks, is a shrub of the family Proteaceae that is native only to eastern New South Wales in Australia. It occurs naturally in woodland, open forest, and heathland on sandstone soi ...
'') and paperbark tea-tree (''
Leptospermum trinervium ''Leptospermum trinervium'', commonly known as flaky-barked tea-tree, slender tea-tree or paperbark tree, is a species of shrub or small tree that is endemic to eastern Australia. It has papery bark that is shed in thin, flaking layers, narrow e ...
'').


Ecology

Conesticks is killed by fire and regenerates afterwards by canopy-stored seedbank. Plants can live up to 60 years in nature. A field study in
Brisbane Water National Park The Brisbane Water National Park is a protected national park that is located in the Central Coast region of New South Wales, in eastern Australia. The national park is situated north of Sydney, west of , and southwest of . Features The ...
found that ''Petrophile pulchella'' had greater reproductive output in areas that had had two short intervals of under seven years between fires, over areas that had had one short interval, and that plants in these latter areas had greater reproductive output than areas with no intervals under seven years between fires. Seeds can germinate up to 700 days after a bushfire, and it is possible that the seedbank could theoretically last up to 90 years between fires. Native bees are possible pollinators of this species.


References

{{Taxonbar, from=Q7179057 Flora of New South Wales Flora of Queensland pulchella Plants described in 1796 Taxa named by Johann Christoph Wendland