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The Persian horned viper (''Pseudocerastes persicus''), known as the Persian horned viper, false horned viper,Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. . and by other common names, is a species of vipers endemic to the Middle East and Asia. Like all other vipers, it is venomous.


Description

Adults average between in total length (body + tail), with a maximum total length of being reported. Females are usually larger than males. These snakes can attain a considerable weight relative to their size, with specimens sometimes exceeding . The head is broad, flat, distinct from the neck and covered with small, imbricate scales. The snout is short and rounded. The nostrils are positioned dorsolaterally and have valves. The
nasal scale In reptiles, the nasal scale refers to the scale that encloses the nostril.Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Malabar, Florida: Krieger Publishing Company. 359 pp. . Sometimes this ...
is unbroken. The rostral scale is small and wide. The eyes are medium in size with vertically elliptical
pupils The pupil is a black hole located in the center of the iris of the eye that allows light to strike the retina.Cassin, B. and Solomon, S. (1990) ''Dictionary of Eye Terminology''. Gainesville, Florida: Triad Publishing Company. It appears black ...
. There are 15-20
interocular scales In snakes, the interorbital scales, or intersupraoculars, are the scales on the top of the head between the plates surmounting the eyesWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . ( ...
and 15-20 circumorbitals. The supraorbital hornlike structure above each eye consists of small, imbricate scales and is also present in juveniles. There are 11-14
supralabials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
and 13-17
sublabials In reptiles, the sublabial scales, also called lower-labials or infralabials, are those scales that border the mouth opening along the lower jaw. They do not include the median scaleWright AH, Wright AA. 1957. ''Handbook of Snakes''. Comstock Publis ...
. 2-4 rows of small scales separate the supralabial scales from the
suboculars In scaled reptiles, the ocular scales are those forming the margin of the eye.Wright AH, Wright AA. 1957. ''Handbook of Snakes''. Comstock Publishing Associates (7th printing, 1985). 1105 pp. . The name originates from the term ''oculus'' which ...
. The body is covered with weakly to strongly keeled dorsal scales. On many of these, the keel terminates before the end of the scale and forms a bump. Many others form a point. At midbody, there are 21-25 scale rows, none of them oblique. There are 134-163 ventral scales and 35-50 paired subcaudals. The tail is short.


Common names

Persian horned viper, false horned viper, Persian horned desert viper,Brown JH. 1973. ''Toxicology and Pharmacology of Venoms from Poisonous Snakes''. Springfield, Illinois: Charles C. Thomas. 184 pp. LCCCN 73-229. . eye-horned viper.U.S. Navy. 1991. ''Poisonous Snakes of the World''. United States Government. New York: Dover Publications Inc. 203 pp. .


Etymology

The species name comes from where it is most usually found, Persia (present-day Iran), and the hornlike structures above its eyes.


Geographic range

''Pseudocerastes persicus'' is found in Iran, Pakistan, Afghanistan, northern Oman, the United Arab Emirates, Jordan, and throughout Mesopotamia. The type locality is listed as ''"Perse"'' (= Persia).


Habitat

This species likes sandy (but not sand dune) or basalt and limestone rock desert and hill country, usually with some vegetation. It tends to avoid areas of human habitation.


Behavior

These snakes are generally rather slow-moving and may employ various methods of locomotion, including sidewinding, serpentine, and rectilinear. They are also terrestrial and almost totally
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
, only being seen during the day or early evening during colder periods. It is not particularly aggressive, but will hiss loudly when disturbed. It is not capable of sinking into the sand vertically like '' Cerastes''. These feed mainly on lizards, small mammals, mice and occasionally small birds and
arthropods Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
, but will also eat dead food.


Reproduction

It is
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
, and sexually mature females lay 11-21 eggs. When produced, these already contain well-developed
embryo An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male spe ...
s, each of which can be as much as in total length. As a result, they hatch after only 30–32 days at 31 °C and then measure in total length. They do well in captivity and are relatively easy to breed.Spawls S, Branch B. 1995. ''The Dangerous Snakes of Africa''. Dubai: Ralph Curtis Books. Oriental Press. 192 pp. .


Venom

''Pseudocerastes persicus'' venom exhibits strong hemorrhagic activity typical of most vipers. No antivenom is available for bites from this species, although it is reported that a polyvalent antiserum does offer some protection.


References


Further reading

* Boulenger GA. 1896. ''Catalogue of the Snakes in the British Museum (Natural History). Volume III., Containing the ... Viperidæ.'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiv + 727 pp. + Plates I.- XXV. (Genus ''Pseudocerastes'' and species ''Pseudocerastes persicus'', p. 501). * Duméril A-M-C, Bibron G, Duméril A A 1854. ''Erpétologie générale ou histoire naturelle complète des reptiles. Tome septième. Deuxième partie. Comprenant l'histoire des serpents venimeux''. (= ''General Herpetology or Complete Natural History of the Reptiles. Volume 7. Second Part. Containing the'' 'Natural'' ''History of the Venomous Snakes''). Paris: Roret. xii + pp. 781–1536. (''Cerastes persicus'', pp. 1443–1444). * Joger U. 1984. ''The venomous snakes of the Near and Middle East''. Wiesbaden: Dr. Ludwig Reichert Verlag. 175 pp. * Latifi M. 1991. ''The Snakes of Iran''. Second Edition. Oxford, Ohio: Published by the Dept. of the Environment and the Society for the Study of Amphibians and Reptiles. 156 pp. . * Marx H, Rabb GB. 1965. Relationships and Zoogeography of the Viperine Snakes (Family Viperidae). ''Fieldiana Zool.'' 44 (21): 162-206. * Mendelssohn H. 1965. On the biology of venomous snakes of Israel. Part II. ''Israeli Journal of Zoology'' 14: 185-212. *Obst FJ. 1983. ''Zur Kenntnis der Schlangengattung'' Vipera. (= On Knowledge of the Snake Genus ''Vipera''). ''Zool. Abh. staatl. Mus. Tierkunde Dresden'' 38: 229-235.


External links

*
Sand viper page
a
Plumed Serpent
Accessed 19 August 2007. {{Taxonbar, from=Q1505139 Viperinae Reptiles of Pakistan Reptiles of the Middle East Reptiles described in 1854 Taxa named by André Marie Constant Duméril Taxa named by Gabriel Bibron Taxa named by Auguste Duméril Snakes of Jordan