The whitenose pigfish (''Perryena leucometopon'') is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

belonging to the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Congiopodidae Congiopodidae, commonly known as pigfishes, horsefishes and racehorses, is a family of ray-finned fish classified with in the order Scorpaeniformes. These fishes are native to the Southern Hemisphere. Taxonomy Congiopodidae was first formally ...

, the horsefishes or pigfishes. It is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to the waters off southern and western Australia. It is the only species in the

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

''Perryena'' and the

classification Classification is a process related to categorization, the process in which ideas and objects are recognized, differentiated and understood. Classification is the grouping of related facts into classes. It may also refer to: Business, organizat ...

of that genus in the family Congiopodidae is not universally agreed upon.

Taxonomy

The whitenose pigfish was first formally described in 1922 as ''Congiopodus leucometopon'' by the British-born Australian

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...

ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...

herpetologist Herpetology (from Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is the branch of zoology concerned with the study of amphibians (including frogs, toads, salamanders, newts, and caecilians (gymnophiona)) and rept ...

, and

ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...

Edgar Ravenswood Waite Edgar Ravenswood Waite (5 May 1866 – 19 January 1928) was a British/Australian zoologist, ichthyologist, herpetologist, and ornithologist. Waite was born in Leeds, Yorkshire, England, the second son of John Waite, a bank clerk, and his wife J ...

with the type locality given as the beach at Glenelg on

Gulf St Vincent Gulf St Vincent, sometimes referred to as St Vincent Gulf, St Vincent's Gulf or Gulf of St Vincent, is the eastern of two large inlets of water on the southern coast of Australia, in the state of South Australia, the other being the larger Sp ...

South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...

. In 1940

Gilbert Percy Whitley Gilbert Percy Whitley (9 June 1903 – 18 July 1975) was a British-born Australian ichthyologist and malacologist who was Curator of Fishes at the Australian Museum in Sydney for about 40 years. He was born at Swaythling, Southampton, England, an ...

reclassified this species in the monotypic genus ''Perryena''. A recent study placed the whitenose pigfish into an expanded stonefish clade, the

Synanceiidae Synanceiinae is a subfamily of venomous ray-finned fishes, waspfishes, which is classified as part of the family Scorpaenidae, the scorpionfishes and their relatives. These fishes are found in the Indo-Pacific oceans. They are primarily marin ...

, because all of these fish have a lachrymal sabre that can project a switch-blade-like mechanism out from underneath their eye. Other classify it within the monogeneric family Perryenidae but the 5th edition of ''

Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...

'' retains this taxon within the family Congiopodidae.

Etymology

The whitenose pigfish was given the specific name ''leucometopon'' by Waite, this is a compound of ''leuco'', which means, "white" and ''metopon'', meaning "forehead", an allusion to the white patch at the anterior part of the head. The genus name of ''Perryena'' was coined by Whitley and honours George Perry, the English naturalist who originally named the genus ''

Congiopodus ''Congiopodus'' is a genus of marine ray-finned fishes belonging to the family Congiopodidae, the pigfishes or horsefishes. These fishes are found in the southern Atlantic and Pacific oceans. Taxonomy Congiopodus was first formally described as ...

'', which Waite had originally classified this species in.

Description

The whitenose pigfish has a dorsal fin which starts over the centre of the eye and which contains 15 spines, with the fourth spine being the longest, and 9 soft rays. Unlike the species in the gneus ''Congiopodus'' there are 3 spines, as opposed to none, in the

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

and 6 or 7 soft rays. There are no scales on the skin. The forehead, first dorsal-fin spine, the anterior part of the snout and the tip of the chin are white, separated from the dark brown remainder of the head and body by a black stripe. This is a small species in which the type specimens were and in length.

Distribution and habitat

The whitenose pigfish is endemic to southern and western Australia where there have been scattered records from the central coast of South Australia and off the southern and western coasts of

Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...

as far north as

Port Denison Port Denison is a town of 3000 residents in the Mid West region of Western Australia. Its local government area is the Shire of Irwin and it is located southwest of Dongara on the Indian Ocean coast. History Port Denison was initially known ...

. It is a reef associated,

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

species found on the

continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...

References

{{Taxonbar, from=Q1662072 Congiopodidae Monotypic fish genera Fish described in 1922 Taxa named by Edgar Ravenswood Waite Taxa named by Gilbert Percy Whitley