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Peppermint (''Mentha'' × ''piperita'') is a
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
species of mint, a cross between
watermint ''Mentha aquatica'' (water mint; syn. ''Mentha hirsuta'' Huds.Euro+Med Plantbase Project''Mentha aquatica'') is a perennial plant, perennial flowering plant in the Mentha, mint family Lamiaceae. It grows in moist places and is native to much of ...
and spearmint. Indigenous to Europe and the Middle East, the plant is now widely spread and cultivated in many regions of the world.Euro+Med Plantbase Project
''Mentha'' × ''piperita''
/ref> It is occasionally found in the wild with its parent species.Flora of NW Europe
''Mentha'' × ''piperita''
Although the genus ''Mentha'' comprises more than 25 species, the one in most common use is peppermint. While Western peppermint is derived from ''Mentha × piperita'', Chinese peppermint, or ''bohe'', is derived from the fresh leaves of ''M. haplocalyx''. ''M. × piperita'' and ''M. haplocalyx'' are both recognized as plant sources of
menthol Menthol is an organic compound, more specifically a monoterpenoid, made synthetically or obtained from the oils of corn mint, peppermint, or other mints. It is a waxy, clear or white crystalline substance, which is solid at room temperature and ...
and menthone, and are among the oldest herbs used for both culinary and medicinal products.


Botany

Peppermint was first described in 1753 by Carl Linnaeus from specimens that had been collected in England; he treated it as a species,Linnaeus, C. (1753). ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
'' 2: 576–577.
but it is now universally agreed to be a hybrid.Harley, R. M. (1975). ''Mentha'' L. In:
Stace, C. A. Clive Anthony Stace (born 1938) is a British botanist and botanical author. He studied at King's College London, graduated from University of London in 1959 and then studied at the Natural History Museum, London. He was awarded a PhD in 1963. ...
, ed. ''Hybridization and the flora of the British Isles'' page 387.
It is a
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
, rhizomatous, perennial plant that grows to be tall, with smooth stems, square in cross section. The
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s are wide-spreading and fleshy, and bear fibrous roots. The
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
can be long and broad. They are dark green with reddish veins, with an acute apex and coarsely toothed margins. The leaves and stems are usually slightly fuzzy. The flowers are purple, long, with a four-lobed
corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name * Corolla (headgear), an ancient headdress in the form of a circlet or crown * ''Corolla'' (gastropod), a genus of moll ...
about diameter; they are produced in whorls (verticillasters) around the stem, forming thick, blunt spikes. Flowering season lasts from mid- to late summer. The chromosome number is variable, with 2n counts of 66, 72, 84, and 120 recorded.Huxley, A., ed. (1992). ''New RHS Dictionary of Gardening''. Macmillan. .Blamey, M. & Grey-Wilson, C. (1989). ''Flora of Britain and Northern Europe''. Peppermint is a fast-growing plant; once it sprouts, it spreads very quickly.


Ecology

Peppermint typically occurs in moist habitats, including stream sides and drainage ditches. Being a hybrid, it is usually
sterile Sterile or sterility may refer to: *Asepsis, a state of being free from biological contaminants * Sterile (archaeology), a sediment deposit which contains no evidence of human activity *Sterilization (microbiology), any process that eliminates or ...
, producing no seeds and reproducing only
vegetatively Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...
, spreading by its runners. Outside of its native range, areas where peppermint was formerly grown for oil often have an abundance of feral plants, and it is considered
invasive Invasive may refer to: *Invasive (medical) procedure *Invasive species *Invasive observation, especially in reference to surveillance *Invasively progressive spread of disease from one organ in the body to another, especially in reference to cancer ...
in Australia, the Galápagos Islands, New Zealand,Pacific Island Ecosystems at Risk
''Mentha'' x ''piperita''
and the United States in the Great Lakes region, noted since 1843.


Cultivation

Peppermint generally grows best in moist, shaded locations, and expands by underground rhizomes. Young shoots are taken from old stocks and dibbled into the ground about 0.5 m (1.5 ft) apart. They grow quickly and cover the ground with runners if it is permanently moist. For the home gardener, it is often grown in containers to restrict rapid spreading. It grows best with a good supply of water, without being water-logged, and planted in areas with partial sun to shade. The leaves and flowering tops are used; they are collected as soon as the flowers begin to open and can be dried. The wild form of the plant is less suitable for this purpose, with cultivated plants having been selected for more and better oil content. They may be allowed to lie and wilt a little before distillation, or they may be taken directly to the still.


Cultivars

Several cultivars have been selected for garden use: * ''Mentha'' × ''piperita'' 'Candymint' has reddish stems. * ''Mentha'' × ''piperita'' 'Chocolate Mint'. Its flowers open from the bottom up; its flavour is reminiscent of the flavour in
Andes Chocolate Mints Andes Chocolate Mints are small rectangular candies. The crème de menthe variety consists of one mint-green layer sandwiched in between two cocoa based chocolate flavored layers. The candies are usually wrapped in green foil and imprinted with ...
, a popular confection. * ''Mentha'' × ''piperita'' 'Citrata' includes a number of varieties including 'eau de Cologne' mint, grapefruit mint, lemon mint, and orange mint. Its leaves are aromatic and hairless. * ''Mentha'' × ''piperita'' 'Crispa' has wrinkled leaves. * ''Mentha'' × ''piperita'' 'Lavender Mint' * ''Mentha'' × ''piperita'' 'Lime Mint' has lime-scented foliage. * ''Mentha'' × ''piperita'' 'Variegata' has mottled green and pale yellow leaves. Commercial cultivars may include: * Dulgo pole * Zefir * Bulgarian population #2 * Clone 11-6-22 * Clone 80-121-33 * Mitcham Digne 38 * Mitcham Ribecourt 19 * '
Todd's Mitcham Peppermint (''Mentha'' × ''piperita'') is a hybrid species of mint, a cross between watermint and spearmint. Indigenous to Europe and the Middle East, the plant is now widely spread and cultivated in many regions of the world.Euro+Med Plantba ...
', a
verticillium wilt Verticillium wilt is a wilt disease affecting over 350 species of eudicot plants. It is caused by six species of ''Verticillium'' fungi: ''V. dahliae'', ''V. albo-atrum'', ''V. longisporum'', ''V. nubilum'', ''V. theobromae'' and ''V. tricorpu ...
-resistant cultivar produced from a breeding and test program of
atomic gardening Atomic gardening is a form of mutation breeding where plants are exposed to radiation. Some of the mutations produced thereby have turned out to be useful. Typically this is gamma radiation in which case it is a produced by cobalt-60. The pra ...
at Brookhaven National Laboratory from the mid-1950s * '
Refined Murray Peppermint (''Mentha'' × ''piperita'') is a hybrid species of mint, a cross between watermint and spearmint. Indigenous to Europe and the Middle East, the plant is now widely spread and cultivated in many regions of the world.Euro+Med Plantbas ...
', also verticillium-resistant * '
Roberts Mitcham Peppermint (''Mentha'' × ''piperita'') is a hybrid species of mint, a cross between watermint and spearmint. Indigenous to Europe and the Middle East, the plant is now widely spread and cultivated in many regions of the world.Euro+Med Plantba ...
', also verticillium-resistant and also the product of mutation breeding


Diseases

Verticillium wilt Verticillium wilt is a wilt disease affecting over 350 species of eudicot plants. It is caused by six species of ''Verticillium'' fungi: ''V. dahliae'', ''V. albo-atrum'', ''V. longisporum'', ''V. nubilum'', ''V. theobromae'' and ''V. tricorpu ...
is a major constraint in peppermint cultivation. 'Todd's Mitcham', 'Refined Murray', 'Roberts Mitcham' (see above), and a few other cultivars have some degree of
resistance Resistance may refer to: Arts, entertainment, and media Comics * Either of two similarly named but otherwise unrelated comic book series, both published by Wildstorm: ** ''Resistance'' (comics), based on the video game of the same title ** ''T ...
.


Production

In 2020, world production of peppermint was 48,437 tonnes, led by Morocco with 83% of the world total and Argentina with 14% (table). In the United States, Oregon and Washington produce most of the country's peppermint, the leaves of which are processed for the
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
to produce flavorings mainly for chewing gum and toothpaste.


Chemical constituents

Peppermint has a high menthol content. The essential oil also contains menthone and carboxyl esters, particularly menthyl acetate. Dried peppermint typically has 0.3–0.4% of volatile oil containing menthol (7–48%), menthone (20–46%), menthyl acetate (3–10%),
menthofuran Menthofuran is an organic compound found in a variety of essential oils including that of pennyroyal (''Mentha pulegium''). It is highly toxic and believed to be the primary toxin in pennyroyal responsible for its potentially fatal effects. Af ...
(1–17%), and 1,8-
cineol Eucalyptol is a monoterpenoid. A colorless liquid, it is a bicyclic ether. Eucalyptol has a fresh mint-like smell and a spicy, cooling taste. It is insoluble in water, but miscible with organic solvents. Eucalyptol makes up ~70% - 90% of eucal ...
(3–6%). Peppermint oil also contains small amounts of many additional compounds, including limonene, pulegone, caryophyllene, and pinene. Peppermint contains terpenoids and
flavonoid Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans. Chemically, flavonoids ...
s such as eriocitrin,
hesperidin Hesperidin is a flavanone glycoside found in citrus fruits. Its aglycone form is called hesperetin. Its name is derived from the word "hesperidium", for fruit produced by citrus trees. Hesperidin was first isolated in 1828 by French chemist M. ...
, and kaempferol 7-O-rutinoside.


Oil

Peppermint oil has a high concentration of natural pesticides, mainly pulegone (found mainly in ''M. arvensis'' var. ''piperascens'' (cornmint, field mint, or Japanese mint), and to a lesser extent (6,530 ppm) in ''Mentha'' × ''piperita'' subsp. ''notho'') and menthone. It is known to repel some pest insects, including mosquitos, and has uses in organic gardening. It is also widely used to repel rodents. Peppermint oil can also be used as an effective remedy for nausea and digestive issues. The chemical composition of the essential oil from peppermint (''Mentha'' × ''piperita'' L.) was analyzed by GC/FID and GC-MS. The main constituents were
menthol Menthol is an organic compound, more specifically a monoterpenoid, made synthetically or obtained from the oils of corn mint, peppermint, or other mints. It is a waxy, clear or white crystalline substance, which is solid at room temperature and ...
(40.7%) and menthone (23.4%). Further components were (±)- menthyl acetate, 1,8- cineole, limonene, beta-pinene, and
beta-caryophyllene Caryophyllene (), more formally (−)-β-caryophyllene, (BCP), is a natural bicyclic sesquiterpene that is a constituent of many essential oils, especially clove oil, the oil from the stems and flowers of '' Syzygium aromaticum'' (cloves), the es ...
.


Research and health effects

Peppermint oil is under preliminary research for its potential as a short-term treatment for irritable bowel syndrome, and has supposed uses in traditional medicine for minor ailments. Peppermint oil and leaves have a cooling effect when used topically for muscle pain,
nerve pain Neuropathic pain is pain caused by damage or disease affecting the somatosensory system. Neuropathic pain may be associated with abnormal sensations called dysesthesia or pain from normally non-painful stimuli (allodynia). It may have continuous ...
, relief from itching, or as a fragrance. High oral doses of peppermint oil (500 mg) can cause mucosal irritation and mimic heartburn. Peppermint roots bioaccumulate radium, so the plant may be effective for phytoremediation of radioactively contaminated soil.


Culinary and other uses

Fresh or dried peppermint leaves are often used alone in peppermint tea or with other herbs in herbal teas (tisanes, infusions). Peppermint is used for flavouring ice cream,
candy Candy, also called sweets (British English) or lollies (Australian English Australian English (AusE, AusEng, AuE, AuEng, en-AU) is the set of varieties of the English language native to Australia. It is the country's common language an ...
,
fruit preserve Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the method ...
s,
alcoholic beverage An alcoholic beverage (also called an alcoholic drink, adult beverage, or a drink) is a drink that contains ethanol, a type of alcohol that acts as a drug and is produced by fermentation of grains, fruits, or other sources of sugar. The c ...
s, chewing gum, toothpaste, and some shampoos, soaps, and skin care products. Menthol activates cold-sensitive TRPM8 receptors in the skin and mucosal tissues, and is the primary source of the cooling sensation that follows the topical application of peppermint oil. File:Candy-Cane-Classic.jpg, Candy canes are one of the most common peppermint-flavored candies File:CSA-Chocolate-Mint.jpg, ''Mentha'' × ''piperita'' hybrid known as 'Chocolate Mint' File:Pfefferminze ies.jpg, Freeze-dried leaves Peppermint oil is also used in construction and plumbing to test for the tightness of pipes and disclose leaks by its odor.


Safety

Medicinal uses of peppermint have not been approved as effective or safe by the US Food and Drug Administration. With caution that the concentration of the peppermint constituent pulegone should not exceed 1% (140 mg), peppermint preparations are considered safe by the
European Medicines Agency The European Medicines Agency (EMA) is an agency of the European Union (EU) in charge of the evaluation and supervision of medicinal products. Prior to 2004, it was known as the European Agency for the Evaluation of Medicinal Products or Euro ...
when used in topical formulations for adult subjects. Diluted peppermint essential oil is safe for oral intake when only a few drops are used. Although peppermint is commonly available as a herbal supplement, no established, consistent manufacturing standards exist for it, and some peppermint products may be contaminated with toxic metals or other substituted compounds. Skin rashes, irritation, or allergic reactions may result from applying peppermint oil to the skin, and its use on the face or chest of young children may cause side effects if the oil menthol is inhaled. A common side effect from oral intake of peppermint oil or capsules is heartburn. Oral use of peppermint products may have adverse effects when used with iron supplements, cyclosporine, medicines for heart conditions or high blood pressure, or medicines to decrease stomach acid.


Standardization

*
ISO 676 ISO is the most common abbreviation for the International Organization for Standardization. ISO or Iso may also refer to: Business and finance * Iso (supermarket), a chain of Danish supermarkets incorporated into the SuperBest chain in 2007 * Is ...
:1995—contains the information about the nomenclature of the variety and cultivars *
ISO 5563 ISO is the most common abbreviation for the International Organization for Standardization. ISO or Iso may also refer to: Business and finance * Iso (supermarket), a chain of Danish supermarkets incorporated into the SuperBest chain in 2007 * Is ...
:1984—a specification for its dried leaves of Mentha piperita Linnaeus * Peppermint oil—
ISO 856 ISO is the most common abbreviation for the International Organization for Standardization. ISO or Iso may also refer to: Business and finance * Iso (supermarket), a chain of Danish supermarkets incorporated into the SuperBest chain in 2007 * Iso ...
:2006


See also

* Eucalyptus * Peppermint extract


References

{{Authority control Antiemetics Flora of Europe Herbs Medicinal plants Mentha Plants described in 1753 Hybrid plants