Pentatomidae is a family of

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...

s belonging to the order

Hemiptera Hemiptera (; ) is an order (biology), order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, Reduviidae, assassin bugs, Cimex, bed bugs, and shield bugs. ...

, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamily

Pentatomoidea The Pentatomoidea are a superfamily of insects in the Heteroptera suborder of the Hemiptera order. As Hemiptera, they share a common arrangement of sucking mouthparts. The roughly 7000 species under Pentatomoidea are divided into 21 families (16 ...

, and contains around 900 genera and over 4700 species.Robert G. Foottit, Peter H. Adler ''Insect Biodiversity: Science and Society'', John Wiley and Sons, 2009, As hemipterans, the pentatomids have piercing sucking mouthparts, and most are

phytophagous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...

, including several species which are severe pests on agricultural

crop A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydropon ...

s. However, some species, particularly in the subfamily

Asopinae Asopinae are a subfamily of stink bugs (family Pentatomidae). They are predatory stink bugs that are useful as biological control agents against pests, even against other Pentatomid species, which are all herbivorous. Genera # '' Afrius'' ( ...

, are predatory and may be considered beneficial.

Etymology

The name "Pentatomidae" is from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''pente'' meaning "five" and ''tomos'' meaning "section", and refers to the five segments of their antennae. Pentatomids are generally called "shield bugs" in British

English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ide ...

, or "stink bugs" in

American English American English, sometimes called United States English or U.S. English, is the set of variety (linguistics), varieties of the English language native to the United States. English is the Languages of the United States, most widely spoken lan ...

. However, the term shield bugs is also applied broadly to include several related families (e.g.

Acanthosomatidae Acanthosomatidae is a family of Hemiptera, commonly named "shield bugs," for which Kumar in his 1979 world revision recognized 47 genera; now this number is 55 genera, with about 200 species, and is one of the least diversified families within P ...

Scutelleridae Scutelleridae is a family of true bugs. They are commonly known as jewel bugs or metallic shield bugs due to their often brilliant coloration. They are also known as shield-backed bugs due to the enlargement of the thoracic scutellum into a con ...

, and

Cydnidae Cydnidae are a family of pentatomoid bugs, known by common names including burrowing bugs or burrower bugs. As the common name would suggest, many members of the group live a subterranean lifestyle, burrowing into soil using their head and forel ...

), or specifically only to refer to species in the family Acanthosomatidae. The term shield bug refers to the generalized body shape of adult bugs in these families which resembles a

heraldic shield In heraldry, an escutcheon () is a shield that forms the main or focal element in an achievement of arms. The word can be used in two related senses. In the first sense, an escutcheon is the shield upon which a coat of arms is displayed. In the s ...

when viewed from above. The American name "stink bug" is specific to the Pentatomidae, and refers to their ability to release a pungent defensive spray when threatened, disturbed, or crushed. The composition of this spray may vary between species, and even by sex or age, but generally includes

aldehyde In organic chemistry, an aldehyde () is an organic compound containing a functional group with the structure . The functional group itself (without the "R" side chain) can be referred to as an aldehyde but can also be classified as a formyl grou ...

s and alkanes. Descriptions of the smells vary widely, and include oily, dusty, woody and earthy, and like coriander. In some species, the liquid contains cyanide compounds and a rancid almond scent, used to protect themselves and discourage predators. The term "stink bug" may also be a vernacular for unrelated insects such as pinacate beetles (in the genus ''

Eleodes ''Eleodes'' is a genus of darkling beetles, family Tenebrionidae. They are commonly known as pinacate beetles or desert stink beetles. They are endemic to western North America, with many species found in the Sonoran Desert. The name ''pinacate ...

'').

Taxonomy

There are several subfamilies, of which the Aphylinae is often given family status, but is here retained as a subfamily, following Grazia ''et al.'' (2008). The subfamilies include: * Aphylinae Bergroth, 1906 – Australia # '' Aphylus'' Bergroth, 1906 # '' Neoaphylum'' Štys & Davidová-Vilímová, 2001 *

Spinola, 1850 ** ''

Troilus Troilus ( or ; grc, Τρωΐλος, Troïlos; la, Troilus) is a legendary character associated with the story of the Trojan War. The first surviving reference to him is in Homer's ''Iliad,'' composed in the late 8th century BCE. In Greek myth ...

'' Stål, 1868 *Cyrtocorinae Distant, 1880 # '' Ceratozygum'' Horváth, 1916 # '' Cyphothyrea'' Horváth, 1916 # '' Cyrtocoris'' White, 1842 # '' Pseudocyrtocoris'' Jensen-Haarup, 1926 *

Discocephalinae The Discocephalinae are a subfamily of shield bugs, erected by Fieber in 1860,Fieber FX (1860) ''Die europäischen Hemiptera. Halbflüger. (Rhynchota Heteroptera).'' 444 pp. Carl Gerold's Sohn (Wien). and found worldwide. Tribes and genera ''B ...

Fieber, 1860 ** '' Discocephala'' Laporte, 1832 ** ''

Ochlerus ''Ochlerus'' is a genus of true bugs belonging to the family Pentatomidae Pentatomidae is a family of insects belonging to the order Hemiptera, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamil ...

'' Spinola, 1837 *Edessinae Fieber, 1860 # '' Anisoedessa'' Nunes & Fernandes, 2019 # '' Brachystethus'' Laporte, 1833 # '' Doesburgedessa'' Fernandes, 2010 # '' Edessa (bug)'' Fabricius, 1803 # '' Grammedessa'' Correia & Fernandes, 2016 # '' Lopadusa'' Stål, 1860 # '' Mediocampus'' Thomas, 1994 # '' Neopharnus'' Van Duzee, 1910 # ''

Olbia (bug) Olbia (, ; sc, Terranoa; sdn, Tarranoa) is a city and communes of Italy, commune of 60,346 inhabitants (May 2018) in the Italy, Italian insular province of Sassari in northeastern Sardinia, Italy, in the historical region of Gallura. Called i ...

'' Stål, 1862 # '' Pantochlora'' Stål, 1870 # '' Paraedessa'' Silva & Fernandes, 2013 # '' Peromatus'' Amyot & Serville, 1843 # '' Pharnus'' Stål, 1867 # '' Plagaedessa'' Almeida & Fernandes, 2018 # '' Platistocoris'' Rider, 1998 # '' Praepharnus'' Barber & Bruner, 1932 *

Pentatominae Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera. Species in this subfamily are phytophages and several of them are consid ...

Leach, 1815 ** '' Bathycoelia'' Amyot & Serville, 1843 ** ''

Pentatoma ''Pentatoma'' is a genus of shield bugs in the family Pentatomidae Pentatomidae is a family of insects belonging to the order Hemiptera, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamily Pen ...

'' Olivier, 1789 *

Phyllocephalinae The Phyllocephalinae are a subfamily of shield bugs erected by Amyot and Serville in 1843. Tribes and genera ''BioLib'' lists the following genera in four tribes:Cressona'' Dallas, 1851 ** '' Megarrhamphus'' Bergroth, 1891 ** '' Phyllocephala'' Laporte, 1833 ** '' Tetroda'' Amyot & Serville, 1843 * Podopinae Amyot & Serville, 1843 ** '' Graphosoma'' Laporte de Castelnau, 1833 ** '' Podops'' Laporte de Castelnau, 1833 *Serbaninae (monotypic) # '' Serbana borneensis'' Distant, 1906 *Stirotarsinae (monotypic) # '' Stirotarsus'' Bergroth, 1911 *''

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...

'' # '' Antillosciocoris'' Thomas, 2005 # ''

Asopus Asopus (; grc, Ἀ̄σωπός ''Āsōpos'') is the name of four different rivers in Greece and one in Turkey. In Greek mythology, it was also the name of the gods of those rivers. Zeus carried off Aegina, Asopus' daughter, and Sisyphus, who h ...

'' Burmeister, 1834 # '' Jostenicoris'' Arnold, 2011

Description

All pentatomids have 5-segmented antennae, and 3 tarsal segments on each foot. They generally have a large triangular scutellum in the center of the back. The body shape of adult pentatomids is generally " shieldlike," when viewed from above, but this varies between species, and is not true for the immature nymphal stages. The forewings of stink bugs are called hemelytra, with the basal half thickened while the apex is membranous. At rest, the wings are laid across the back of the insect, with the membranous wingtips overlapping. The hindwings are entirely membranous.

Economics

Several stink bugs and shield bugs are considered agricultural pests, because they can grow into large populations that feed on crops, damaging production, and they are resistant to many pesticides. They are a threat to cotton, corn, sorghum, soybeans, native and ornamental trees, shrubs, vines, weeds, and many cultivated crops. In

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

, some species of stink bugs are called ''jumil'', ''chinche de monte'', ''xotlinilli'', or ''chumil'' (e.g. '' Edessa mexicana''). They are most often eaten in the states of Morelos and

Guerrero Guerrero is one of the 32 states that comprise the 32 Federal Entities of Mexico. It is divided in 81 municipalities and its capital city is Chilpancingo and its largest city is Acapulcocopied from article, GuerreroAs of 2020, Guerrero the pop ...

. The flavor is sometimes said to resemble cinnamon, or sometimes a bitter medicinal flavor. ''Jumiles'' may be used for making sauces or as

taco A taco (, , ) is a traditional Mexican food consisting of a small hand-sized corn- or wheat-based tortilla topped with a filling. The tortilla is then folded around the filling and eaten by hand. A taco can be made with a variety of fillin ...

filling. Since recent arrival in the U.S., populations of the

brown marmorated stink bug The brown marmorated stink bug (''Halyomorpha halys'') is an insect in the family Pentatomidae, native to China, Japan, Korea and other Asian regions. In September 1998 it was collected in Allentown, Pennsylvania, where it is believed to have be ...

have grown significantly. As of October 2014, brown marmorated stink bugs can be found in 41 out of 50 states within the U.S. In 2016 New Zealand's MPI put out an alert to prevent this invasive species from entering via imported cargo. Acoloba lanceolata01

European species

European species within this family include: * ''

Acrosternum arabicum ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * '' Acrosternum arabicum'' Wagner, 1959 * '' Acrost ...

'' Wagner, 1959 * ''

Acrosternum heegeri ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' Fieber, 1861 * ''

Acrosternum malickyi ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' Josifov & Heiss, 1989 * ''

Acrosternum millierei ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' (Mulsant & Rey, 1866) * ''

Acrosternum rubescens ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' (Noualhier, 1893) * '' Aelia acuminata'' (Linnaeus, 1758) * '' Aelia albovittata'' Fieber, 1868 * ''

Aelia angusta Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Stehlik, 1976 * ''

Aelia cognata Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Fieber, 1868 * ''

Aelia cribrosa Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Fieber, 1868 * ''

Aelia furcula Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Fieber, 1868 * '' Aelia germari'' Kuster, 1852 * '' Aelia klugii'' Hahn, 1833 * '' Aelia notata'' Rey, 1887 * ''

Aelia rostrata Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Boheman, 1852 * '' Aelia sibirica'' Reuter, 1884 * ''

Aelia virgata Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' (Herrich-Schäffer, 1841) * '' Ancyrosoma leucogrammes'' (Gmelin, 1790) * '' Andrallus spinidens'' (Fabricius, 1787) * '' Antheminia absinthii'' (Wagner, 1952) * ''

Antheminia aliena ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the g ...

'' (Reuter, 1891) * '' Antheminia lunulata'' (Goeze, 1778) * '' Antheminia pusio'' (Kolenati, 1846) * '' Antheminia varicornis'' (Jakovlev, 1874) * ''

Apodiphus amygdali ''Apodiphus amygdali'' is a species of shield bug belonging to the family Pentatomidae, subfamily Pentatominae. Distribution This species can be found in Austria, Albania, Bulgaria, Greece, Italy and North Macedonia. Description ''Apodiphus am ...

'' (Germar, 1817) * '' Arma custos'' (Fabricius, 1794) * '' Arma insperata'' Horvath, 1899 * '' Asaroticus solskyi'' Jakovlev, 1873 * '' Bagrada abeillei'' Puton, 1881 * '' Bagrada confusa'' Horvath, 1936 * '' Bagrada elegans'' Puton, 1873 * '' Bagrada funerea'' Horvath, 1901 * '' Bagrada hilaris'' (Burmeister, 1835) * '' Bagrada stolida'' (Herrich-Schäffer, 1839) * '' Bagrada turcica'' Horvath, 1936 * '' Brachynema cinctum'' (Fabricius, 1775) * '' Brachynema germarii'' (Kolenati, 1846) * ''

Brachynema purpureomarginatum ''Brachynema'' is the scientific name of two genera of organisms and may refer to: *Brachynema (bug), ''Brachynema'' (bug), a genus of stinkbugs in the family Pentatomidae *Brachynema (plant), ''Brachynema'' (plant), a genus of plants in the fam ...

'' (Rambur, 1839) * '' Capnoda batesoni'' Jakovlev, 1889 * '' Carpocoris coreanus'' Distant, 1899 * '' Carpocoris fuscispinus'' (Boheman, 1850) * '' Carpocoris melanocerus'' (Mulsant & Rey, 1852) * '' Carpocoris pudicus'' (Poda, 1761) * '' Carpocoris purpureipennis'' (De Geer, 1773) * '' Chlorochroa juniperina'' (Linnaeus, 1758) * '' Chlorochroa pinicola'' (Mulsant & Rey, 1852) * '' Chlorochroa reuteriana'' (Kirkaldy, 1909) * '' Chroantha ornatula'' (Herrich-Schäffer, 1842) * '' Codophila varia'' (Fabricius, 1787) * '' Crypsinus angustatus'' (Baerensprung, 1859) * '' Derula flavoguttata'' Mulsant & Rey, 1856 * '' Dolycoris baccarum'' (Linnaeus, 1758) * ''

Dolycoris numidicus ''Dolycoris'' is a genus of shield bug in the family Pentatomidae. Species * ''Dolycoris alobatus'' Hasan & Afzal, 1990 * ''Dolycoris baccarum'' (Linnaeus, 1758) - Sloe Bug * ''Dolycoris bengalensis'' Zaidi, 1995 * ''Dolycoris brachyserratus' ...

'' Horvath, 1908 * '' Dryadocoris apicalis'' (Herrich-Schäffer, 1842) * '' Dybowskyia reticulata'' (Dallas, 1851) * ''

Dyroderes umbraculatus ''Dyroderes umbraculatus'' is a shield bug belonging to the family Pentatomidae, subfamily Pentatominae. The species was first described by Johan Christian Fabricius in 1775. The genus ''Dyroderes'' has one living representative, but ''BioLib' ...

'' (Fabricius, 1775) * '' Eudolycoris alluaudi'' (Noualhier, 1893) * '' Eurydema cyanea'' (Fieber, 1864) * '' Eurydema dominulus'' (Scopoli, 1763) * ''

Eurydema eckerleini ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * ''Eurydema herbacea'' (Her ...

'' Josifov, 1961 * '' Eurydema fieberi'' Schummel, 1837 * '' Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' (Herrich-Schäffer, 1833) * '' Eurydema lundbaldi'' Lindberg, 1960 * '' Eurydema maracandica'' Oshanin, 1871 * '' Eurydema nana'' Fuente, 1971 * '' Eurydema oleracea'' (Linnaeus, 1758) * '' Eurydema ornata'' (Linnaeus, 1758) * '' Eurydema rotundicollis'' (Dohrn, 1860) * ''

Eurydema rugulosa ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (f ...

'' (Dohrn, 1860) * '' Eurydema sea'' Pericart & De la Rosa 2004 * '' Eurydema spectabilis'' Horvath, 1882 * '' Eurydema ventralis'' Kolenati, 1846 * '' Eysarcoris aeneus'' (Scopoli, 1763) * '' Eysarcoris ventralis'' (Westwood, 1837) * '' Eysarcoris venustissimus'' (Schrank, 1776) * '' Graphosoma interruptum'' White, 1839 * ''

Graphosoma italicum ''Graphosoma italicum'' is a species of shield bug in the family Pentatomidae. It is also known as the Striped bug (or Italian striped bug) and Minstrel bug. Description ''G. italicum'' can reach a length of . The body is almost round, with a l ...

'' (Müller, 1766) * '' Graphosoma lineatum'' (Linnaeus, 1758) * '' Graphosoma melanoxanthum'' Horvath, 1903 * '' Graphosoma semipunctatum'' (Fabricius, 1775) * ''

'' (Stål, 1855) * '' Holcogaster fibulata'' (Germar, 1831) * '' Holcostethus albipes'' (Fabricius, 1781) * ''

Holcostethus evae ''Holcostethus'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae Pentatominae is a subfamily of Pentatomidae Pentatomidae is a family of insects belonging to the order Hemiptera, generally called shi ...

'' Ribes, 1988 * ''

Holcostethus sphacelatus ''Holcostethus'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. Species These 18 species belong to the genus ''Holcostethus''. * '' Holcostethus abbreviatus'' Uhler, 1872 * '' Holcostethus albipes'' (Fabr ...

'' (Fabricius, 1794) * '' Jalla dumosa'' (Linnaeus, 1758) * '' Leprosoma inconspicuum'' Baerensprung, 1859 * '' Leprosoma stali'' Douglas & Scott, 1868 * '' Leprosoma tuberculatum'' Jakovlev, 1874 * '' Macrorhaphis acuta'' Dallas, 1851 * '' Mecidea lindbergi'' Wagner, 1954 * '' Mecidea pallidissima'' Jensen-Haarup, 1922 * '' Menaccarus arenicola'' (Scholz, 1847) * '' Menaccarus deserticola'' Jakovlev, 1900 * '' Menaccarus dohrnianus'' (Mulsant & Rey, 1866) * '' Menaccarus turolensis'' Fuente, 1971 * '' Mustha spinosula'' (Lefèbvre, 1831) * '' Neostrachia bisignata'' (Walker, 1867) * '' Neottiglossa bifida'' (A. Costa, 1847) * '' Neottiglossa flavomarginata'' (Lucas, 1849) * ''

Neottiglossa leporina ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * ''Neottiglossa bifida'' (A.Costa, 1847) * ''Neottig ...

'' (Herrich-Schäffer, 1830) * '' Neottiglossa lineolata'' (Mulsant & Rey, 1852) * '' Neottiglossa pusilla'' (Gmelin, 1790) * ''

Nezara viridula ''Nezara viridula'', commonly known as the southern green stink bug (USA), southern green shield bug (UK) or green vegetable bug (Australia and New Zealand), is a plant-feeding stink bug. Believed to have originated in Ethiopia, it can now be fo ...

'' (Linnaeus, 1758) * '' Palomena formosa'' Vidal, 1940 * '' Palomena prasina'' (Linnaeus, 1761) * '' Palomena viridissima'' (Poda, 1761) * ''

Pentatoma rufipes The forest bug or red-legged shieldbug (''Pentatoma rufipes'') is a species of shield bug in the family Pentatomidae, commonly found in most of Europe. It inhabits forests, woodlands, orchards, and gardens. Description ''P. rufipes'' can reach a ...

'' (Linnaeus, 1758) * '' Peribalus congenitus'' Putshkov, 1965 * '' Peribalus inclusus'' (Dohrn, 1860) * '' Peribalus strictus'' (Fabricius, 1803) * '' Perillus bioculatus'' (Fabricius, 1775) * ''

Picromerus bidens ''Picromerus bidens'', the spiny shieldbug or spiked shieldbug, is a carnivorous species of shield bug in the family Pentatomidae. Distribution and habitat This species has a wide distribution in the Palaearctic, from 64°N to North Africa and ...

'' (Linnaeus, 1758) * '' Picromerus brachypterus'' Ahmad & Onder, 1990 * '' Picromerus conformis'' (Herrich-Schäffer, 1841) * ''

Picromerus nigridens ''Picromerus'' is a genus of shield bugs in the family Pentatomidae. Genera * ''Picromerus bidens'' (Linnaeus, 1758) * ''Picromerus brachypterus'' Ahmad & Önder, 1990 * ''Picromerus conformis'' (Herrich-Schäffer, 1841) * ''Picromerus elevat ...

'' (Fabricius, 1803) * ''

Piezodorus lituratus ''Piezodorus lituratus'', the gorse shield bug, is a species of Pentatomidae, a family of shield bugs. Varieties *''Piezodorus lituratus var. lituratus'' (Johann Christian Fabricius, Fabricius, 1794) *''Piezodorus lituratus var. alliaceus'' (Ger ...

'' (Fabricius, 1794) * ''

Piezodorus punctipes ''Piezodorus'' is a genus of Pentatomidae, a family of shield bugs. Species *''Piezodorus guildinii'' Westwood *''Piezodorus lituratus'' (Fabricius, 1794) – gorse shield bug *''Piezodorus punctipes'' Puton, 1889 *''Piezodorus rubrofasciatus' ...

'' Puton, 1889 * ''

Piezodorus teretipes ''Piezodorus'' is a genus of Pentatomidae, a family of shield bugs. Species *'' Piezodorus guildinii'' Westwood *''Piezodorus lituratus'' (Fabricius, 1794) – gorse shield bug *''Piezodorus punctipes'' Puton, 1889 *''Piezodorus rubrofasciatus ...

'' (Stål, 1865) * '' Pinthaeus sanguinipes'' (Fabricius, 1781) * '' Podops annulicornis'' Jakovlev, 1877 * '' Podops calligerus'' Horvath, 1887 * '' Podops curvidens'' Costa, 1843 * '' Podops dilatatus'' Puton, 1873 * '' Podops inunctus'' (Fabricius, 1775) * '' Podops rectidens'' Horvath, 1883 * '' Putonia torrida'' Stål, 1872 * ''

Rhacognathus punctatus ''Rhacognathus punctatus'', the Heather Bug, is a species of stink bugs (family Pentatomidae). Description ''Rhacognathus punctatus'' can reach a length of . This predatory stink bug has a mottled metallic bronze-brown shieldbug, extended should ...

'' (Linnaeus, 1758) * '' Rhaphigaster nebulosa'' (Poda, 1761) * ''

Rubiconia intermedia ''Rubiconia'' is a genus of true bugs belonging to the family Pentatomidae. The species of this genus are found in Europe and Japan. Species: * ''Rubiconia intermedia'' (Wolff, 1811) References

{{Taxonbar, from=Q4048584 Pentatomidae ...

'' (Wolff, 1811) * '' Schyzops aegyptiaca'' (Lefèbvre, 1831) * '' Sciocoris angularis'' Puton, 1889 * ''

Sciocoris angusticollis ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Puton, 1895 * '' Sciocoris conspurcatus'' Klug, 1845 * '' Sciocoris convexiusculus'' Puton, 1874 * '' Sciocoris cursitans'' (Fabricius, 1794) * ''

Sciocoris deltocephalus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1861 * '' Sciocoris distinctus'' Fieber, 1851 * '' Sciocoris galiberti'' Ribaut, 1926 * ''

Sciocoris helferi ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1851 * ''

Sciocoris hoberlandti ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Wagner, 1954 * '' Sciocoris homalonotus'' Fieber, 1851 * '' Sciocoris luteolus'' Fieber, 1861 * '' Sciocoris macrocephalus'' Fieber, 1851 * '' Sciocoris maculatus'' Fieber, 1851 * '' Sciocoris microphthalmus'' Flor, 1860 * '' Sciocoris modestus'' Horvath, 1903 * ''

Sciocoris ochraceus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1861 * ''

Sciocoris orientalis ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * ''Sciocoris cerrutii'' Wa ...

'' Linnavuori, 1960 * '' Sciocoris pallens'' Klug, 1845 * '' Sciocoris pentheri'' Wagner, 1953 * '' Sciocoris pictus'' Wagner, 1959 * '' Sciocoris sideritidis'' Wollaston, 1858 * '' Sciocoris sulcatus'' Fieber, 1851 * '' Sciocoris umbrinus'' (Wolff, 1804) * '' Sciocoriscanariensis'' Lindberg, 1953 * '' Scotinophara sicula'' (A. Costa, 1841) * '' Scotinophara subalpina'' (Bergroth, 1893) * '' Stagonomus amoenus'' (Brullé, 1832) * '' Stagonomus bipunctatus'' (Linnaeus, 1758) * '' Stagonomus devius'' Seidenstucker, 1965 * '' Stagonomus grenieri'' (Signoret, 1865) * '' Staria lunata'' (Hahn, 1835) * '' Stenozygum coloratum'' (Klug, 1845) * '' Sternodontus binodulus'' Jakovlev, 1893 * '' Sternodontus obtusus'' Mulsant & Rey, 1856 * '' Tarisa dimidiatipes'' Puton, 1874 * '' Tarisa elevata'' Reuter, 1901 * '' Tarisa flavescens'' Amyot & Serville, 1843 * '' Tarisa pallescens'' Jakovlev, 1871 * '' Tarisa salsolae'' Kerzhner, 1964 * '' Tarisa subspinosa'' (Germar, 1839) * '' Tholagmus flavolineatus'' (Fabricius, 1798) * '' Tholagmus strigatus'' (Herrich-Schäffer, 1835) * '' Trochiscocoris hemipterus'' (Jakovlev, 1879) * '' Trochiscocoris rotundatus'' Horvath, 1895 * ''

Troilus luridus ''Troilus luridus'', also known as the bronze shieldbug, is a species of Pentatomidae, shield bug found in Europe. It was Species description, first described by the Danish zoology, zoologist Johan Christian Fabricius in 1775. Shieldbugs are gene ...

'' (Fabricius, 1775) * '' Ventocoris achivus'' (Horvath, 1889) * '' Ventocoris falcatus'' (Cyrillus, 1791) * '' Ventocoris fischeri'' (Herrich-Schäffer, 1851) * '' Ventocoris halophilum'' (Jakovlev, 1874) * '' Ventocoris modestus'' (Jakovlev, 1880) * '' Ventocoris philalyssum'' (Kiritshenko, 1916) * '' Ventocoris ramburi'' (Horvath, 1908) * '' Ventocoris rusticus'' (Fabricius, 1781) * '' Ventocoris trigonus'' (Krynicki, 1871) * '' Vilpianus galii'' (Wolff, 1802) * '' Zicrona caerulea'' (Linnaeus, 1758) Eurydema oleracea.jpg, Image:Chinche hedionda depredadora (Euthyrhynchus floridanus).jpg, '' Euthyrhynchus floridanus'', nymph Image:Troilus luridus 20020828.jpg, ''

'' Image:Pentatomidae spec.jpg, '' Raphigaster nebulosa'' Image:Eurydema ornatum en Tenerife.jpg, '' Eurydema ornatum'' File:Perillus nymph.jpg, '' Perillus'', nymph

References

External links

When Twenty-Six Thousand Stinkbugs Invade Your Home
by

Kathryn Schulz Kathryn Schulz is an American journalist and author. She is a staff writer at ''The New Yorker''. In 2016, she won the Pulitzer Prize for Feature Writing for her article on the risk of a major earthquake and tsunami in the Pacific Northwest. B ...

. ''The New Yorker,'' March 12, 2018.

Ed Yong Edmund Soon-Weng Yong (born 17 December 1981) is a British-American science journalist. He is a staff member at '' The Atlantic'', which he joined in 2015. In 2021 he received a Pulitzer Prize for Explanatory Reporting for a series on the COVI ...

called it a "pure delight."
I.A.D. Robertson: ''The Pentatomoidea (Hemiptera: Heteroptera) of Sub-Saharan Africa : a database''
Malindi, 2009

– photos and information

* I. A. D. Robertson: ''The Pentatomoidea (Hemiptera: Heteroptera) of Sub-Saharan Africa : a database''. Malindi, 2009
Online version in Repository Naturalis Leiden
{{Taxonbar, from=Q652142 Heteroptera families Shield bugs Taxa named by William Elford Leach la:Cimex foetidus