Pentatomidae is a family of

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

s belonging to the order

Hemiptera Hemiptera (; ) is an order (biology), order of insects, commonly called true bugs, comprising over 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, Reduviidae, assassin bugs, Cimex, bed bugs, and shield bugs. ...

, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamily Pentatomoidea, and contains around 900 genera and over 4700 species.Robert G. Foottit, Peter H. Adler ''Insect Biodiversity: Science and Society'', John Wiley and Sons, 2009, As hemipterans, the pentatomids have piercing sucking mouthparts, and most are phytophagous, including several species which are severe pests on

agricultural Agriculture or farming is the practice of cultivating Plant, plants and livestock. Agriculture was the key development in the rise of Sedentism, sedentary human civilization, whereby farming of Domestication, domesticated species created food ...

crop A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydroponic ...

s. However, some species, particularly in the subfamily Asopinae, are predatory and may be considered beneficial.

Etymology

The name "Pentatomidae" is from the Greek ''pente'' meaning "five" and ''tomos'' meaning "section", and refers to the five segments of their antennae. Pentatomids are generally called "shield bugs" in British

English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ide ...

, or "stink bugs" in

American English American English, sometimes called United States English or U.S. English, is the set of variety (linguistics), varieties of the English language native to the United States. English is the Languages of the United States, most widely spoken lan ...

. However, the term shield bugs is also applied broadly to include several related families (e.g.

Acanthosomatidae Acanthosomatidae is a family of Hemiptera, commonly named "shield bugs," for which Kumar in his 1979 world revision recognized 47 genera; now this number is 55 genera, with about 200 species, and is one of the least diversified families within P ...

Scutelleridae Scutelleridae is a family of true bugs. They are commonly known as jewel bugs or metallic shield bugs due to their often brilliant coloration. They are also known as shield-backed bugs due to the enlargement of the thoracic scutellum into a con ...

, and Cydnidae), or specifically only to refer to species in the family Acanthosomatidae. The term shield bug refers to the generalized body shape of adult bugs in these families which resembles a heraldic shield when viewed from above. The American name "stink bug" is specific to the Pentatomidae, and refers to their ability to release a pungent defensive spray when threatened, disturbed, or crushed. The composition of this spray may vary between species, and even by sex or age, but generally includes

aldehyde In organic chemistry, an aldehyde () is an organic compound containing a functional group with the structure . The functional group itself (without the "R" side chain) can be referred to as an aldehyde but can also be classified as a formyl grou ...

s and

alkane In organic chemistry, an alkane, or paraffin (a historical trivial name that also has other meanings), is an acyclic saturated hydrocarbon. In other words, an alkane consists of hydrogen and carbon atoms arranged in a tree structure in which ...

s. Descriptions of the smells vary widely, and include oily, dusty, woody and earthy, and like

coriander Coriander (;

. In some species, the liquid contains cyanide compounds and a rancid almond scent, used to protect themselves and discourage predators. The term "stink bug" may also be a vernacular for unrelated insects such as pinacate beetles (in the genus ''

Eleodes ''Eleodes'' is a genus of darkling beetles, family Tenebrionidae. They are commonly known as pinacate beetles or desert stink beetles. They are endemic to western North America, with many species found in the Sonoran Desert. The name ''pinacate ...

'').

Taxonomy

There are several subfamilies, of which the Aphylinae is often given family status, but is here retained as a subfamily, following Grazia ''et al.'' (2008). The subfamilies include: * Aphylinae Bergroth, 1906 – Australia # '' Aphylus'' Bergroth, 1906 # '' Neoaphylum'' Štys & Davidová-Vilímová, 2001 * Asopinae Spinola, 1850 ** '' Troilus'' Stål, 1868 *Cyrtocorinae Distant, 1880 # '' Ceratozygum'' Horváth, 1916 # '' Cyphothyrea'' Horváth, 1916 # '' Cyrtocoris'' White, 1842 # '' Pseudocyrtocoris'' Jensen-Haarup, 1926 *

Discocephalinae The Discocephalinae are a subfamily of shield bugs, erected by Fieber in 1860,Fieber FX (1860) ''Die europäischen Hemiptera. Halbflüger. (Rhynchota Heteroptera).'' 444 pp. Carl Gerold's Sohn (Wien). and found worldwide. Tribes and genera ''Bi ...

Fieber, 1860 ** '' Discocephala'' Laporte, 1832 ** ''

Ochlerus ''Ochlerus'' is a genus of true bugs belonging to the family Pentatomidae Pentatomidae is a family of insects belonging to the order Hemiptera, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamil ...

'' Spinola, 1837 *Edessinae Fieber, 1860 # '' Anisoedessa'' Nunes & Fernandes, 2019 # '' Brachystethus'' Laporte, 1833 # '' Doesburgedessa'' Fernandes, 2010 # '' Edessa (bug)'' Fabricius, 1803 # '' Grammedessa'' Correia & Fernandes, 2016 # '' Lopadusa'' Stål, 1860 # '' Mediocampus'' Thomas, 1994 # '' Neopharnus'' Van Duzee, 1910 # ''

Olbia (bug) Olbia (, ; sc, Terranoa; sdn, Tarranoa) is a city and communes of Italy, commune of 60,346 inhabitants (May 2018) in the Italy, Italian insular province of Sassari in northeastern Sardinia, Italy, in the historical region of Gallura. Called i ...

'' Stål, 1862 # '' Pantochlora'' Stål, 1870 # '' Paraedessa'' Silva & Fernandes, 2013 # '' Peromatus'' Amyot & Serville, 1843 # '' Pharnus'' Stål, 1867 # '' Plagaedessa'' Almeida & Fernandes, 2018 # '' Platistocoris'' Rider, 1998 # '' Praepharnus'' Barber & Bruner, 1932 *

Pentatominae Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera. Species in this subfamily are phytophages and several of them are consid ...

Leach, 1815 ** ''

Bathycoelia ''Bathycoelia''Amyot CJ-B, Audinet-Serville JG (1843) ''Histoire naturelle des insectes. Hémiptères. Librairie encyclopédique de Roret'', Paris. XXIII, 110. is a genus of shield bugs in the subfamily Pentatominae and the monotypic tribe Bathyc ...

'' Amyot & Serville, 1843 ** ''

Pentatoma ''Pentatoma'' is a genus of shield bugs in the family Pentatomidae Pentatomidae is a family of insects belonging to the order Hemiptera, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamily Pen ...

'' Olivier, 1789 * Phyllocephalinae Amyot & Serville, 1843 ** ''

Cressona ''Cressona''Dallas WS (1851) ''List Specimens Hem. Ins. Coll. Brit. Museum'' 1. is a genus of shield bugs in the subfamily Phyllocephalinae, erected by William Sweetland Dallas in 1851. Species The following are included in ''BioLib.cz'': # ''Cr ...

'' Dallas, 1851 ** ''

Megarrhamphus ''Megarrhamphus'' is a genus of Asian Pentatomidae, shield bugs in the subfamily Phyllocephalinae and typical of the Tribe (biology), tribe Megarrhamphini, erected by Ernst Evald Bergroth in 1891.Bergroth EE (1891) ''Rev. Ent. Français'', 10. S ...

'' Bergroth, 1891 ** '' Phyllocephala'' Laporte, 1833 ** '' Tetroda'' Amyot & Serville, 1843 *

Podopinae Podopinae, known as turtle bugs, are a subfamily of the insect family Pentatomidae. The type genus is ''Podops''. Tribes and Genera ''BioLib'' lists: Brachycerocorini Auth. Davidova-Vilimova & Stys, 1993 # '' Bolbocoris'' Amyot & Serville, 1843 ...

Amyot & Serville, 1843 ** ''

Graphosoma __NOTOC__ ''Graphosoma'' is a genus bugs in the family Pentatomidae. They are commonly known as striped shield bugs for their distinctive markings. Species Subgenus ''Graphosoma'' *'' Graphosoma alkani'' Lodos, 1959 *'' Graphosoma consimile'' Ho ...

'' Laporte de Castelnau, 1833 ** ''

Podops ''Podops''Laporte de Castelnau FL (1833) Mémoire de cinquante espèces nouvelles ou peu connues d'Insectes. ''Annales de la Société entomologique de France'' 1: 386–415. is a genus of Palaearctic bugs, in the family Pentatomidae; it is the ...

'' Laporte de Castelnau, 1833 *Serbaninae (monotypic) # ''

Serbana The Serbaninae are a highly modified subfamily of Pentatomidae, containing only a single known species, ''Serbana borneensis''. It is known from very few specimens, from Malaysia and Borneo, and was originally described in the family Phloeidae, ...

borneensis'' Distant, 1906 *Stirotarsinae (monotypic) # '' Stirotarsus'' Bergroth, 1911 *'' incertae sedis'' # '' Antillosciocoris'' Thomas, 2005 # '' Asopus'' Burmeister, 1834 # '' Jostenicoris'' Arnold, 2011

Description

All pentatomids have 5-segmented antennae, and 3 tarsal segments on each foot. They generally have a large triangular scutellum in the center of the back. The body shape of adult pentatomids is generally " shieldlike," when viewed from above, but this varies between species, and is not true for the immature nymphal stages. The forewings of stink bugs are called

hemelytra An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alterna ...

, with the basal half thickened while the apex is membranous. At rest, the wings are laid across the back of the insect, with the membranous wingtips overlapping. The hindwings are entirely membranous.

Economics

Several stink bugs and shield bugs are considered agricultural

pest Pest or The Pest may refer to: Science and medicine * Pest (organism), an animal or plant deemed to be detrimental to humans or human concerns ** Weed, a plant considered undesirable * Infectious disease, an illness resulting from an infection ** ...

s, because they can grow into large populations that feed on crops, damaging production, and they are resistant to many

pesticide Pesticides are substances that are meant to control pests. This includes herbicide, insecticide, nematicide, molluscicide, piscicide, avicide, rodenticide, bactericide, insect repellent, animal repellent, microbicide, fungicide, and lampri ...

s. They are a threat to cotton, corn, sorghum, soybeans, native and ornamental trees, shrubs, vines, weeds, and many cultivated crops. In

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

, some species of stink bugs are called ''jumil'', ''chinche de monte'', ''xotlinilli'', or ''chumil'' (e.g. '' Edessa mexicana''). They are most often eaten in the states of

Morelos Morelos (), officially the Free and Sovereign State of Morelos ( es, Estado Libre y Soberano de Morelos), is one of the 32 states which comprise the Federal Entities of Mexico. It is divided into 36 municipalities and its capital city is Cuer ...

and Guerrero. The flavor is sometimes said to resemble cinnamon, or sometimes a bitter medicinal flavor. ''Jumiles'' may be used for making sauces or as taco filling. Since recent arrival in the U.S., populations of the

brown marmorated stink bug The brown marmorated stink bug (''Halyomorpha halys'') is an insect in the family Pentatomidae, native to China, Japan, Korea and other Asian regions. In September 1998 it was collected in Allentown, Pennsylvania, where it is believed to have be ...

have grown significantly. As of October 2014, brown marmorated stink bugs can be found in 41 out of 50 states within the U.S. In 2016 New Zealand's MPI put out an alert to prevent this invasive species from entering via imported cargo. Acoloba lanceolata01

European species

European species within this family include: * ''

Acrosternum arabicum ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * '' Acrosternum arabicum'' Wagner, 1959 * '' Acrost ...

'' Wagner, 1959 * ''

Acrosternum heegeri ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' Fieber, 1861 * ''

Acrosternum malickyi ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' Josifov & Heiss, 1989 * ''

Acrosternum millierei ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' (Mulsant & Rey, 1866) * ''

Acrosternum rubescens ''Acrosternum'' is a genus of stink bugs in the family Pentatomidae. There are more than 20 described species in ''Acrosternum''. Species These 23 species belong to the genus ''Acrosternum'': * ''Acrosternum arabicum'' Wagner, 1959 * '' Acroste ...

'' (Noualhier, 1893) * ''

Aelia acuminata ''Aelia acuminata'', common name Bishop's Mitre, is a species of ''shield bug'' belonging to the family Pentatomidae. Distribution This species is present in most of Europe, in North Africa and in Northern Asia (excluding China). Habitat These ...

'' (Linnaeus, 1758) * '' Aelia albovittata'' Fieber, 1868 * ''

Aelia angusta Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Stehlik, 1976 * ''

Aelia cognata Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Fieber, 1868 * ''

Aelia cribrosa Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Fieber, 1868 * ''

Aelia furcula Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Fieber, 1868 * '' Aelia germari'' Kuster, 1852 * '' Aelia klugii'' Hahn, 1833 * '' Aelia notata'' Rey, 1887 * ''

Aelia rostrata Aelia may refer to: * Aelia (bug), ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens), a plebeian family at Rome, which flourished from the fifth century BC until at least the third c ...

'' Boheman, 1852 * ''

Aelia sibirica Aelia may refer to: * ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens) The gens Aelia, occasionally written Ailia, was a plebeian family in Rome, which flourished from the fifth century BC until ...

'' Reuter, 1884 * ''

Aelia virgata Aelia may refer to: * ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae * Aelia (gens) The gens Aelia, occasionally written Ailia, was a plebeian family in Rome, which flourished from the fifth century BC until ...

'' (Herrich-Schäffer, 1841) * '' Ancyrosoma leucogrammes'' (Gmelin, 1790) * '' Andrallus spinidens'' (Fabricius, 1787) * '' Antheminia absinthii'' (Wagner, 1952) * ''

Antheminia aliena ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the g ...

'' (Reuter, 1891) * ''

Antheminia lunulata ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the ...

'' (Goeze, 1778) * ''

Antheminia pusio ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the ...

'' (Kolenati, 1846) * ''

Antheminia varicornis ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the ...

'' (Jakovlev, 1874) * ''

Apodiphus amygdali ''Apodiphus amygdali'' is a species of shield bug belonging to the family Pentatomidae, subfamily Pentatominae. Distribution This species can be found in Austria, Albania, Bulgaria, Greece, Italy and North Macedonia. Description ''Apodiphus am ...

'' (Germar, 1817) * '' Arma custos'' (Fabricius, 1794) * '' Arma insperata'' Horvath, 1899 * '' Asaroticus solskyi'' Jakovlev, 1873 * ''

Bagrada abeillei ''Bagrada'' is a genus of stink bugs in the family Pentatomidae. Species * '' Bagrada abeillei'' Puton, 1881 * ''Bagrada elegans'' Puton, 1873 * '' Bagrada funerea'' Horvath, 1901 * ''Bagrada hilaris ''Bagrada hilaris'' is a species of shield ...

'' Puton, 1881 * '' Bagrada confusa'' Horvath, 1936 * ''

Bagrada elegans ''Bagrada'' is a genus of stink bugs in the family Pentatomidae Pentatomidae is a family of insects belonging to the order Hemiptera, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamily Pentatomo ...

'' Puton, 1873 * ''

Bagrada funerea ''Bagrada'' is a genus of stink bugs in the family Pentatomidae. Species * ''Bagrada abeillei'' Puton, 1881 * ''Bagrada elegans'' Puton, 1873 * '' Bagrada funerea'' Horvath, 1901 * ''Bagrada hilaris ''Bagrada hilaris'' is a species of shield b ...

'' Horvath, 1901 * ''

Bagrada hilaris ''Bagrada hilaris'' is a species of shield bug known by the common names bagrada bug and painted bug. It could be mistaken for or erroneously referred to as harlequin bug. It is native to southeastern Africa. It is known elsewhere as an introduce ...

'' (Burmeister, 1835) * ''

Bagrada stolida ''Bagrada'' is a genus of stink bugs in the family Pentatomidae. Species * ''Bagrada abeillei'' Puton, 1881 * ''Bagrada elegans'' Puton, 1873 * ''Bagrada funerea'' Horvath, 1901 * ''Bagrada hilaris'' (Burmeister, 1835) (Bagrada bug) * ''Bagrada ...

'' (Herrich-Schäffer, 1839) * ''

Bagrada turcica ''Bagrada'' is a genus of stink bugs in the family Pentatomidae. Species * ''Bagrada abeillei'' Puton, 1881 * ''Bagrada elegans'' Puton, 1873 * ''Bagrada funerea'' Horvath, 1901 * ''Bagrada hilaris'' (Burmeister, 1835) (Bagrada bug) * ''Bagrada ...

'' Horvath, 1936 * '' Brachynema cinctum'' (Fabricius, 1775) * '' Brachynema germarii'' (Kolenati, 1846) * ''

Brachynema purpureomarginatum ''Brachynema'' is the scientific name of two genera of organisms and may refer to: *Brachynema (bug), ''Brachynema'' (bug), a genus of stinkbugs in the family Pentatomidae *Brachynema (plant), ''Brachynema'' (plant), a genus of plants in the fam ...

'' (Rambur, 1839) * '' Capnoda batesoni'' Jakovlev, 1889 * ''

Carpocoris coreanus ''Carpocoris'' is a genus of shield bug in the family Pentatomidae. The species of this genus are quite similar.Carpocoris fuscispinus ''Carpocoris fuscispinus'' is a species of shield bug in the family Pentatomidae. Subspecies * ''Carpocoris fuscispinus hahni'' (Flor, 1856) * ''Carpocoris fuscispinus incerta'' (Tamanini, 1959) * ''Carpocoris fuscispinus maculosa'' (Tamanini, ...

'' (Boheman, 1850) * ''

Carpocoris melanocerus ''Carpocoris'' is a genus of shield bug in the family Pentatomidae. The species of this genus are quite similar.Carpocoris pudicus ''Carpocoris pudicus'' is a species of shield bug in the family Pentatomidae. Distribution This species is widespread in most of central and southern Europe (Albania, Austria, Bulgaria, Croatia, Czech Republic, France, Germany, Greece, Italy, No ...

'' (Poda, 1761) * ''

Carpocoris purpureipennis ''Carpocoris purpureipennis'' is a species of shield bug of the family Pentatomidae, subfamily Pentatominae. Subspecies * ''Carpocoris purpureipennis purpureipennis'' (De Geer, 1773) * ''Carpocoris purpureipennis sexmaculatus'' Péneau, 1921 ...

'' (De Geer, 1773) * ''

Chlorochroa juniperina ''Chlorochroa juniperina'' is a European species of shield bug in the tribe Nezarini Nezarini is a tribe of stink bugs in the family Pentatomidae. Genera There are at least 21 described genera in the Nezarini including: * ''Acrosternum ...

'' (Linnaeus, 1758) * ''

Chlorochroa pinicola ''Chlorochroa pinicola'' is a European species of shield bug in the tribe Nezarini Nezarini is a tribe of stink bugs in the family Pentatomidae. Genera There are at least 21 described genera in the Nezarini including: * ''Acrosternum ' ...

'' (Mulsant & Rey, 1852) * ''

Chlorochroa reuteriana ''Chlorochroa''Stål C (1872) ''K. svenska VetenskAkad. Handl.'' 10, no. 4. is a genus of shield (stink) bugs in the family Pentatomidae, found in Europe and North America. There are over 20 described species in ''Chlorochroa''. Description A ...

'' (Kirkaldy, 1909) * '' Chroantha ornatula'' (Herrich-Schäffer, 1842) * '' Codophila varia'' (Fabricius, 1787) * ''

Crypsinus angustatus ''Crypsinus'' is a monotypic genus of Pentatomidae, shield bugs in the Tribe (biology), tribe Graphosomatini,BioLib.cz
genus '' ...

'' (Baerensprung, 1859) * '' Derula flavoguttata'' Mulsant & Rey, 1856 * ''

Dolycoris baccarum ''Dolycoris baccarum'', the sloe bug or hairy shieldbug, is a species of shield bug in the family Pentatomidae. Distribution and habitat This species is widespread in most of Europe and Central Asia. These shield bugs mainly inhabit hedgerows ...

'' (Linnaeus, 1758) * ''

Dolycoris numidicus ''Dolycoris'' is a genus of shield bug in the family Pentatomidae. Species * ''Dolycoris alobatus'' Hasan & Afzal, 1990 * ''Dolycoris baccarum'' (Linnaeus, 1758) - Sloe Bug * ''Dolycoris bengalensis'' Zaidi, 1995 * ''Dolycoris brachyserratus' ...

'' Horvath, 1908 * ''

Dryadocoris apicalis ''Dryadocoris apicalis'' is a species of shield bugs of the family Pentatomidae. Description ''Dryadocoris apicalis'' can reach an adult length of . Distribution This species can be found in France, Italy, Spain, Portugal, Canary Islands, Moroc ...

'' (Herrich-Schäffer, 1842) * '' Dybowskyia reticulata'' (Dallas, 1851) * ''

Dyroderes umbraculatus ''Dyroderes umbraculatus'' is a shield bug belonging to the family Pentatomidae, subfamily Pentatominae. The species was first described by Johan Christian Fabricius in 1775. The genus ''Dyroderes'' has one living representative, but ''BioLib' ...

'' (Fabricius, 1775) * '' Eudolycoris alluaudi'' (Noualhier, 1893) * ''

Eurydema cyanea ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * '' Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' ...

'' (Fieber, 1864) * '' Eurydema dominulus'' (Scopoli, 1763) * '' Eurydema eckerleini'' Josifov, 1961 * '' Eurydema fieberi'' Schummel, 1837 * ''

Eurydema gebleri ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * '' Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' (H ...

'' Kolenati, 1846 * '' Eurydema herbacea'' (Herrich-Schäffer, 1833) * '' Eurydema lundbaldi'' Lindberg, 1960 * ''

Eurydema maracandica ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' (He ...

'' Oshanin, 1871 * ''

Eurydema nana ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' (He ...

'' Fuente, 1971 * '' Eurydema oleracea'' (Linnaeus, 1758) * ''

Eurydema ornata ''Eurydema ornata'' is a species of shield bug in the family Pentatomidae. Description ''Eurydema ornata'' can reach a length of . The body has gray or bright red colors, with black markings. Sometimes it shows the red patches coloured white an ...

'' (Linnaeus, 1758) * '' Eurydema rotundicollis'' (Dohrn, 1860) * ''

Eurydema rugulosa ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' (He ...

'' (Dohrn, 1860) * '' Eurydema sea'' Pericart & De la Rosa 2004 * ''

Eurydema spectabilis ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' (He ...

'' Horvath, 1882 * ''

Eurydema ventralis ''Eurydema ventralis'' is a shield bug of the family Pentatomidae, subfamily Pentatominae. The species was first described by Friedrich August Rudolph Kolenati in 1846. Description ''Eurydema ventralis'' has a length of about . It typically has ...

'' Kolenati, 1846 * ''

Eysarcoris aeneus ''Eysarcoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. Species * ''Eysarcoris aeneus'' (Scopoli, 1763) * ''Eysarcoris confusus'' Fuente, 1971 * ''Eysarcoris distinctus'' (Schouteden) * ''Eysarcoris ...

'' (Scopoli, 1763) * ''

Eysarcoris ventralis ''Eysarcoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. Species * ''Eysarcoris aeneus'' (Scopoli, 1763) * ''Eysarcoris confusus'' Fuente, 1971 * ''Eysarcoris distinctus'' (Schouteden) * ''Eysarcoris ...

'' (Westwood, 1837) * '' Eysarcoris venustissimus'' (Schrank, 1776) * ''

Graphosoma interruptum __NOTOC__ ''Graphosoma'' is a genus bugs in the family Pentatomidae. They are commonly known as striped shield bugs for their distinctive markings. Species Subgenus ''Graphosoma'' *'' Graphosoma alkani'' Lodos, 1959 *'' Graphosoma consimile'' Ho ...

'' White, 1839 * '' Graphosoma italicum'' (Müller, 1766) * ''

Graphosoma lineatum ''Graphosoma lineatum'' is a species of shield bug in the family Pentatomidae. Description ''G. lineatum'' can reach a length of . The body is almost round, with a large shield. The basic color of the upperside of the body is orange yellow, with ...

'' (Linnaeus, 1758) * ''

Graphosoma melanoxanthum __NOTOC__ ''Graphosoma'' is a genus bugs in the family Pentatomidae. They are commonly known as striped shield bugs for their distinctive markings. Species Subgenus ''Graphosoma'' *''Graphosoma alkani'' Lodos, 1959 *''Graphosoma consimile'' Horv ...

'' Horvath, 1903 * ''

Graphosoma semipunctatum ''Graphosoma semipunctatum'' is a species of true bug living exclusively in the Mediterranean region. The form living on the island of Crete Crete ( el, Κρήτη, translit=, Modern: , Ancient: ) is the largest and most populous of the G ...

'' (Fabricius, 1775) * ''

'' (Stål, 1855) * '' Holcogaster fibulata'' (Germar, 1831) * '' Holcostethus albipes'' (Fabricius, 1781) * '' Holcostethus evae'' Ribes, 1988 * '' Holcostethus sphacelatus'' (Fabricius, 1794) * '' Jalla dumosa'' (Linnaeus, 1758) * '' Leprosoma inconspicuum'' Baerensprung, 1859 * '' Leprosoma stali'' Douglas & Scott, 1868 * '' Leprosoma tuberculatum'' Jakovlev, 1874 * '' Macrorhaphis acuta'' Dallas, 1851 * '' Mecidea lindbergi'' Wagner, 1954 * '' Mecidea pallidissima'' Jensen-Haarup, 1922 * '' Menaccarus arenicola'' (Scholz, 1847) * '' Menaccarus deserticola'' Jakovlev, 1900 * '' Menaccarus dohrnianus'' (Mulsant & Rey, 1866) * '' Menaccarus turolensis'' Fuente, 1971 * '' Mustha spinosula'' (Lefèbvre, 1831) * '' Neostrachia bisignata'' (Walker, 1867) * ''

Neottiglossa bifida ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * '' Neottiglossa bifida'' (A.Costa, 1847) * '' Neott ...

'' (A. Costa, 1847) * ''

Neottiglossa flavomarginata ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * ''Neottiglossa bifida'' (A.Costa, 1847) * '' Neotti ...

'' (Lucas, 1849) * '' Neottiglossa leporina'' (Herrich-Schäffer, 1830) * ''

Neottiglossa lineolata ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * ''Neottiglossa bifida'' (A.Costa, 1847) * '' Neotti ...

'' (Mulsant & Rey, 1852) * ''

Neottiglossa pusilla ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * ''Neottiglossa bifida'' (A.Costa, 1847) * '' Neotti ...

'' (Gmelin, 1790) * '' Nezara viridula'' (Linnaeus, 1758) * '' Palomena formosa'' Vidal, 1940 * ''

Palomena prasina The green shield bug (''Palomena prasina'') is a European shield bug species in the family Pentatomidae. The name might equally apply to several other species in the tribe Nezarini, or if referred-to as a "green stink bug", it might more approp ...

'' (Linnaeus, 1761) * '' Palomena viridissima'' (Poda, 1761) * ''

Pentatoma rufipes The forest bug or red-legged shieldbug (''Pentatoma rufipes'') is a species of shield bug in the family Pentatomidae, commonly found in most of Europe. It inhabits forests, woodlands, orchards, and gardens. Description ''P. rufipes'' can reach a ...

'' (Linnaeus, 1758) * '' Peribalus congenitus'' Putshkov, 1965 * '' Peribalus inclusus'' (Dohrn, 1860) * ''

Peribalus strictus ''Peribalus strictus'', common name Vernal Shieldbug, is a species of shield bugs in the family Pentatomidae. Subspecies *''Peribalus strictus capitatus'' Jakovlev, 1889 *''Peribalus strictus strictus'' (Fabricius, 1803) *''Peribalus strict ...

'' (Fabricius, 1803) * ''

Perillus bioculatus ''Perillus bioculatus'', the two-spotted stink bug or double-eyed soldier bug, is a species of insect in the family Pentatomidae. They are native to North America but have been introduced to Eastern Europe and North India. Both the larval and adul ...

'' (Fabricius, 1775) * '' Picromerus bidens'' (Linnaeus, 1758) * '' Picromerus brachypterus'' Ahmad & Onder, 1990 * '' Picromerus conformis'' (Herrich-Schäffer, 1841) * ''

Picromerus nigridens ''Picromerus'' is a genus of shield bugs in the family Pentatomidae. Genera * ''Picromerus bidens'' (Linnaeus, 1758) * ''Picromerus brachypterus'' Ahmad & Önder, 1990 * ''Picromerus conformis'' (Herrich-Schäffer, 1841) * ''Picromerus elevat ...

'' (Fabricius, 1803) * '' Piezodorus lituratus'' (Fabricius, 1794) * '' Piezodorus punctipes'' Puton, 1889 * ''

Piezodorus teretipes ''Piezodorus'' is a genus of Pentatomidae, a family of shield bugs. Species *'' Piezodorus guildinii'' Westwood *''Piezodorus lituratus'' (Fabricius, 1794) – gorse shield bug *''Piezodorus punctipes'' Puton, 1889 *''Piezodorus rubrofasciatus ...

'' (Stål, 1865) * '' Pinthaeus sanguinipes'' (Fabricius, 1781) * ''

Podops annulicornis ''Podops''Laporte de Castelnau FL (1833) Mémoire de cinquante espèces nouvelles ou peu connues d'Insectes. ''Annales de la Société entomologique de France'' 1: 386–415. is a genus of Palaearctic bugs, in the family Pentatomidae; it is the ...

'' Jakovlev, 1877 * ''

Podops calligerus ''Podops''Laporte de Castelnau FL (1833) Mémoire de cinquante espèces nouvelles ou peu connues d'Insectes. ''Annales de la Société entomologique de France'' 1: 386–415. is a genus of Palaearctic bugs, in the family Pentatomidae; it is the ...

'' Horvath, 1887 * ''

Podops curvidens ''Podops''Laporte de Castelnau FL (1833) Mémoire de cinquante espèces nouvelles ou peu connues d'Insectes. ''Annales de la Société entomologique de France'' 1: 386–415. is a genus of Palaearctic bugs, in the family Pentatomidae; it is the ...

'' Costa, 1843 * '' Podops dilatatus'' Puton, 1873 * ''

Podops inunctus ''Podops''Laporte de Castelnau FL (1833) Mémoire de cinquante espèces nouvelles ou peu connues d'Insectes. ''Annales de la Société entomologique de France'' 1: 386–415. is a genus of Palaearctic bugs, in the family Pentatomidae; it is the ...

'' (Fabricius, 1775) * ''

Podops rectidens ''Podops''Laporte de Castelnau FL (1833) Mémoire de cinquante espèces nouvelles ou peu connues d'Insectes. ''Annales de la Société entomologique de France'' 1: 386–415. is a genus of Palaearctic bugs, in the family Pentatomidae; it is the ...

'' Horvath, 1883 * '' Putonia torrida'' Stål, 1872 * '' Rhacognathus punctatus'' (Linnaeus, 1758) * '' Rhaphigaster nebulosa'' (Poda, 1761) * ''

Rubiconia intermedia ''Rubiconia'' is a genus of true bugs belonging to the family Pentatomidae. The species of this genus are found in Europe and Japan. Species: * ''Rubiconia intermedia'' (Wolff, 1811) References

{{Taxonbar, from=Q4048584 Pentatomidae ...

'' (Wolff, 1811) * '' Schyzops aegyptiaca'' (Lefèbvre, 1831) * '' Sciocoris angularis'' Puton, 1889 * ''

Sciocoris angusticollis ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Puton, 1895 * '' Sciocoris conspurcatus'' Klug, 1845 * '' Sciocoris convexiusculus'' Puton, 1874 * ''

Sciocoris cursitans ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' (Fabricius, 1794) * ''

Sciocoris deltocephalus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1861 * ''

Sciocoris distinctus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Fieber, 1851 * '' Sciocoris galiberti'' Ribaut, 1926 * ''

Sciocoris helferi ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1851 * ''

Sciocoris hoberlandti ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Wagner, 1954 * '' Sciocoris homalonotus'' Fieber, 1851 * ''

Sciocoris luteolus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1861 * ''

Sciocoris macrocephalus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Fieber, 1851 * '' Sciocoris maculatus'' Fieber, 1851 * '' Sciocoris microphthalmus'' Flor, 1860 * '' Sciocoris modestus'' Horvath, 1903 * ''

Sciocoris ochraceus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1861 * ''

Sciocoris orientalis ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * ''Sciocoris cerrutii'' Wa ...

'' Linnavuori, 1960 * ''

Sciocoris pallens ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Klug, 1845 * '' Sciocoris pentheri'' Wagner, 1953 * '' Sciocoris pictus'' Wagner, 1959 * '' Sciocoris sideritidis'' Wollaston, 1858 * ''

Sciocoris sulcatus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' W ...

'' Fieber, 1851 * ''

Sciocoris umbrinus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' (Wolff, 1804) * '' Sciocoriscanariensis'' Lindberg, 1953 * '' Scotinophara sicula'' (A. Costa, 1841) * '' Scotinophara subalpina'' (Bergroth, 1893) * ''

Stagonomus amoenus ''Stagonomus'' is a genus of shieldbug belonging to the family Pentatomidae, subfamily Pentatominae Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 spe ...

'' (Brullé, 1832) * ''

Stagonomus bipunctatus ''Stagonomus'' is a genus of shieldbug belonging to the family Pentatomidae, subfamily Pentatominae. Species Species within this genus include: * ''Stagonomus amoenus'' (Brullé, 1832) * '' Stagonomus bipunctatus'' (Linnaeus, 1758) * ''Stago ...

'' (Linnaeus, 1758) * ''

Stagonomus devius ''Stagonomus'' is a genus of shieldbug belonging to the family Pentatomidae, subfamily Pentatominae. Species Species within this genus include: * ''Stagonomus amoenus'' (Brullé, 1832) * '' Stagonomus bipunctatus'' (Linnaeus, 1758) * '' Stag ...

'' Seidenstucker, 1965 * ''

Stagonomus grenieri ''Stagonomus'' is a genus of shieldbug belonging to the family Pentatomidae, subfamily Pentatominae. Species Species within this genus include: * ''Stagonomus amoenus'' (Brullé, 1832) * '' Stagonomus bipunctatus'' (Linnaeus, 1758) * '' Stag ...

'' (Signoret, 1865) * '' Staria lunata'' (Hahn, 1835) * '' Stenozygum coloratum'' (Klug, 1845) * '' Sternodontus binodulus'' Jakovlev, 1893 * '' Sternodontus obtusus'' Mulsant & Rey, 1856 * '' Tarisa dimidiatipes'' Puton, 1874 * '' Tarisa elevata'' Reuter, 1901 * '' Tarisa flavescens'' Amyot & Serville, 1843 * '' Tarisa pallescens'' Jakovlev, 1871 * '' Tarisa salsolae'' Kerzhner, 1964 * '' Tarisa subspinosa'' (Germar, 1839) * '' Tholagmus flavolineatus'' (Fabricius, 1798) * '' Tholagmus strigatus'' (Herrich-Schäffer, 1835) * '' Trochiscocoris hemipterus'' (Jakovlev, 1879) * '' Trochiscocoris rotundatus'' Horvath, 1895 * ''

Troilus luridus ''Troilus luridus'', also known as the bronze shieldbug, is a species of Pentatomidae, shield bug found in Europe. It was Species description, first described by the Danish zoology, zoologist Johan Christian Fabricius in 1775. Shieldbugs are gene ...

'' (Fabricius, 1775) * '' Ventocoris achivus'' (Horvath, 1889) * '' Ventocoris falcatus'' (Cyrillus, 1791) * '' Ventocoris fischeri'' (Herrich-Schäffer, 1851) * '' Ventocoris halophilum'' (Jakovlev, 1874) * '' Ventocoris modestus'' (Jakovlev, 1880) * '' Ventocoris philalyssum'' (Kiritshenko, 1916) * '' Ventocoris ramburi'' (Horvath, 1908) * '' Ventocoris rusticus'' (Fabricius, 1781) * '' Ventocoris trigonus'' (Krynicki, 1871) * '' Vilpianus galii'' (Wolff, 1802) * ''

Zicrona caerulea ''Zicrona caerulea'', the blue shieldbug, is a species of bug of the family Pentatomidae. Description ''Zicrona caerulea'' can reach an adult size of about . The body is uniformly metallic blue-green (hence the Latin name ''caerulea'', meaning ...

'' (Linnaeus, 1758) Eurydema oleracea.jpg, Image:Chinche hedionda depredadora (Euthyrhynchus floridanus).jpg, '' Euthyrhynchus floridanus'', nymph Image:Troilus luridus 20020828.jpg, ''

'' Image:Pentatomidae spec.jpg, '' Raphigaster nebulosa'' Image:Eurydema ornatum en Tenerife.jpg, ''

Eurydema ornatum ''Eurydema ornata'' is a species of shield bug in the family Pentatomidae. Description ''Eurydema ornata'' can reach a length of . The body has gray or bright red colors, with black markings. Sometimes it shows the red patches coloured white and ...

'' File:Perillus nymph.jpg, ''

Perillus ''Perillus'' is a genus of predatory stink bugs in the family Pentatomidae. There are about seven described species in ''Perillus''. Species These seven species belong to the genus ''Perillus'': * '' Perillus bioculatus'' (Fabricius, 1775) (tw ...

'', nymph

References

External links

When Twenty-Six Thousand Stinkbugs Invade Your Home
by Kathryn Schulz. ''The New Yorker,'' March 12, 2018. Ed Yong called it a "pure delight."
I.A.D. Robertson: ''The Pentatomoidea (Hemiptera: Heteroptera) of Sub-Saharan Africa : a database''
Malindi, 2009

– photos and information

* I. A. D. Robertson: ''The Pentatomoidea (Hemiptera: Heteroptera) of Sub-Saharan Africa : a database''. Malindi, 2009
Online version in Repository Naturalis Leiden
{{Taxonbar, from=Q652142 Heteroptera families Shield bugs Taxa named by William Elford Leach la:Cimex foetidus

Etymology

Taxonomy

Description

Economics

See also

European species

References

External links