Pentacrinites
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''Pentacrinites'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which are ...
s that lived from the
Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...
to the
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
of
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
,
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
. Their stems are pentagonal to star-shaped in cross-section and are the most commonly preserved parts. ''Pentacrinites'' are commonly found in the ''Pentacrinites'' Bed of the
Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
(Lower
Lias Lias may refer to: Geology * Lias Formation, a geologic formation in France *Lias Group, a lithostratigraphic unit in western Europe * Early Jurassic, an epoch People * Godfrey Lias, British author * Mohd Shamsudin Lias (born 1953), Malaysian ...
) of
Lyme Regis Lyme Regis is a town in west Dorset, England, west of Dorchester and east of Exeter. Sometimes dubbed the "Pearl of Dorset", it lies by the English Channel at the Dorset–Devon border. It has noted fossils in cliffs and beaches on the Herita ...
, Dorset, England. ''Pentacrinites'' can be recognized by the extensions (or cirri) all around the stem, which are long, unbranching, and of increasing length further down, the very small cup and 5 long freely branching arms.


Description

Like most
echinoderm An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea ...
s, ''Pentacrinites'' was composed of numerous
calcite Calcite is a Carbonate minerals, carbonate mineral and the most stable Polymorphism (materials science), polymorph of calcium carbonate (CaCO3). It is a very common mineral, particularly as a component of limestone. Calcite defines hardness 3 on ...
plates which were arranged into different body parts. ''Pentacrinites'' had 3 kinds of body parts: arms, cup ( calyx or theca) and stem. The stem consisted of a stack of numerous 5-sided beads (or columnal plates) with a canal at their centre. The stem had flexible appendages (or cirri) that were used to attach an individual. These cirri themselves were connected to specialized columnals called nodals, leaving oval scars after breaking off. The cirri consisted of diamond-shaped plates with a central canal, less flatted further from the stem. The cup-shaped calyx was very small and consisted of two bands of five plates. These were the bases of the five arms. The top of the calyx was covered by numerous small polygonal plates and the mouth and anus were found on this surface. The arms divide frequently, like tree branches, so that at the top end there could be over 50 branches in all. The arms were formed of piles of calcite plates. The arms carried many thin feeding branches (or pinnae, like a fern frond). These pinnae had tube feet, that were covered in mucus, reached into the water and caught plankton. These arms were not very mobile. The arms plates of the arms have an insertion, that formed a grove that ran along the length of the arm and onto the calyx. This served to transport the food particles to the mouth.


Taxonomy

''Pentacrinites'' may have evolved from early, free living isocrinids, which occur today on the sea floor.


Reassigned species

* ''P. caputmedusae'' = ''
Cenocrinus asterius ''Cenocrinus'' is a monotypic genus of stalked crinoids in the family Isselicrinidae. The great West Indian sea lily (''Cenocrinus asterius'') is the only species in the genus and is found in deep waters in the Caribbean Sea and Gulf of Mexico. ...
'' * ''P. subteres'' = '' Balanocrinus subteres'' * ''P. subteroides'' = '' Balanocrinus subteroides''


Ecology

''Pentacrinites'' is a floating sea lily that was attached to driftwood during its adult life. This pseudo-
plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...
ic
lifestyle Lifestyle often refers to: * Lifestyle (sociology), the way a person lives * ''Otium'', ancient Roman concept of a lifestyle * Style of life (german: Lebensstil, link=no), dealing with the dynamics of personality Lifestyle may also refer to: Bus ...
enabled it to exploit food unavailable to other crinoids.


Distribution

''Pentacrinites'' became extinct and has left no direct living descendants. Its remains have been found in many different locations, suggesting it occurred throughout the world's oceans. Free living isocrinids however, still populate the seas, usually at over 150m deep. * ''P. doreckae'' is known from the Lower Jurassic (
Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (My ...
and
Sinemurian In the geologic timescale, the Sinemurian is an age and stage in the Early or Lower Jurassic Epoch or Series. It spans the time between 199.3 ± 2 Ma and 190.8 ± 1.5 Ma (million years ago). The Sinemurian is preceded by the Hettangian and is ...
). * ''P. fossilis'' occurs in the Lower Jurassic (
Sinemurian In the geologic timescale, the Sinemurian is an age and stage in the Early or Lower Jurassic Epoch or Series. It spans the time between 199.3 ± 2 Ma and 190.8 ± 1.5 Ma (million years ago). The Sinemurian is preceded by the Hettangian and is ...
). * ''P. dichotomus'' has been found in the Lower Jurassic (
Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...
). * ''P. dargniesi'' was collected from the Middle Jurassic (
Bajocian In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.3 Ma to around 168.3 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonian Age. Stratig ...
and
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
).


References


Sources

* ''Fossils'' (Smithsonian Handbooks) by David Ward (Page 172)


External links


''Pentacrinites''
in the
Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) originated in the NCEAS-funded Phanerozoic Marine Pale ...
*http://www.fossilmuseum.net/fossils/Crinoids/Pentacrinus-fossilis/Pentacrinus.htm {{Taxonbar, from=Q3899226 Isocrinida Prehistoric crinoid genera Jurassic crinoids Prehistoric echinoderms of Asia Prehistoric echinoderms of Europe Prehistoric echinoderms of North America Sinemurian first appearances Bathonian genus extinctions