Penstemon Ternatus

''Penstemon'' , the beardtongues, is a large genus of roughly 250 species of flowering plants native mostly to the Nearctic, but with a few species also found in the North American portion of the Neotropics. It is the largest genus of flowering plants endemic to North America. Formerly placed in the family Scrophulariaceae by the Cronquist system, new genetic research has placed it in the vastly expanded family Plantaginaceae.

They have opposite leaves, partly tube-shaped, and two-lipped flowers and seed capsules. The most distinctive feature of the genus is the prominent staminode, an infertile stamen. The staminode takes a variety of forms in the different species; while typically a long straight filament extending to the mouth of the corolla, some are longer and extremely hairy, giving the general appearance of an open mouth with a fuzzy tongue protruding and inspiring the common name beardtongue.

Most penstemons are deciduous or semi-evergreen perennials, the remainder being shrubs or subshrubs. Heights can range from 10 cm to as much as 3 metres.

The one Asiatic species previously treated in ''Penstemon'' is now placed in a separate genus ''Pennellianthus''. This leaves ''Penstemon'' a mostly Nearctic genus, with a few neotropical species. Although widespread across North America, and found in habitats ranging from open desert to moist forests, and up to the alpine zone, they are not typically common within their range.

History

John Mitchell published the first scientific description in 1748; although he only named it as ''Penstemon'', we can identify it as '' P. laevigatus''. Linnaeus then included it in his 1753 publication, as ''Chelone pentstemon'', altering the spelling to better correspond to the notion that the name referred to the unusual fifth stamen (Greek "penta-", five). Mitchell's work was reprinted in 1769, continuing with his original spelling, and this was ultimately accepted as the official form, although ''Pentstemon'' continued in use into the 20th century.

Although several more species were found in the 18th century, they continued to be classified in '' Chelone'' until about the 1820s. The period of 1810 to 1850 increased the number of known species from 4 to 63, as expeditions traveled through Mexico and the western United States, followed by another 100 up to 1900.

During this time, seeds began to be offered for sale in Europe, the earliest known dating from 1813, with John Fraser offering four species in London, followed by Flanagan & Nutting offering nine species in their 1835 catalog. Subsequently many hybrids were developed in Europe.

Fieldwork in the remote parts of the Great Basin during the 20th century brought the total number of species known to over 250. The genus was extensively revised by David Keck between 1932 and 1957.

In 1946 the American Penstemon Society (link at bottom) was formed to promote both horticultural and botanical interest.

Uses

Some Native American tribes used penstemons as medicinal remedies for humans and animals.Escamilla, G., et al
''Penstemon ambiguus''.
Medicinal Plants of the Southwest. New Mexico State University. 2001. Updated 2008. Today its primary use is ornamental.

Horticulture

Although penstemons are among the most attractive native flowers of North America, Europe has traditionally been far more active in their hybridization with hundreds of hybrids developed since the early 19th century. The earliest development is somewhat shrouded in mystery; for instance Flanagan & Nutting's 1835 catalog mentions a 'Penstemon Hybridum' but does not describe it.

By 1860, a half-dozen French growers are known to have developed hybrids, most notably Victor Lemoine, while in 1857 the German Wilhelm Pfitzer listed 24 varieties. In 1861 the British Royal Horticultural Society held trials in which 78 varieties were entered. The Scottish firm of John Forbes first offered penstemons in 1870, eventually becoming the biggest grower in the world; in 1884 their catalog listed 180 varieties. By 1900 Forbes had offered 550 varieties, while Lemoine had developed nearly 470 by the time of his death in 1911. Few of these have survived to the present day.

A number of different species have been used in the hybridization process, notably '' P. cobaea'' and '' P. hartwegii''.

In North America, penstemons are often used in xeriscape landscaping, as many are native to desert or alpine regions and thus quite hardy. One of the largest collections of penstemons in North America is found at The Arboretum at Flagstaff, Arizona, which hosts a Penstemon Festival each summer.

Cultivars

The following species and cultivars have gained the Royal Horticultural Society's Award of Garden Merit:-

* 'Andenken an Friedrich Hahn' (deep red)

* 'Beech Park' (pink/white)

* 'Connie's Pink' (rose pink)
* 'Evelyn' (rose pink)
* 'George Home' (red/white)
* 'Hewell Pink Bedder' (pink/white)
* ' Hidcote Pink'
* 'Margery Fish' (purple/blue)
* 'Maurice Gibbs' (purple-red/white)
* 'Osprey' (pink/white)
* ''P. hartwegii'' (scarlet)
* ''P. isophyllus'' (pale pink)
* ''P. pinifolius'' 'Wisley Flame' (orange-red)
* ''P. rupicola'' (pink)

* 'Port Wine' (deep red/white)
* 'Raven' (purple/white)
* 'Rich Ruby'
* 'Roy Davidson' (pink/white)
* 'Rubicundus' (red/white)
* 'Schoenholzeri' (red)
* 'Sour Grapes' (purple/blue)
* 'Stapleford Gem' (purple/blue)

Others include 'Dark Towers', developed by Dale Lindgren at the University of Nebraska.

Species

* '' Penstemon abietinus'' Pennell – Firleaf beardtongue
* '' Penstemon absarokensis'' Evert – Absaroka Range beardtongue
* ''Penstemon acaulis'' L.O. Williams – Stemless beardtongue
* '' Penstemon acuminatus'' Dougl. ex Lindl. – Sharpleaf penstemon
* ''Penstemon alamosensis'' Pennell & Nisbet – Los Alamos beardtongue
* '' Penstemon albertinus'' Greene – Alberta beardtongue
* ''Penstemon albidus'' Nutt. – White penstemon
* ''Penstemon albomarginalis'' M. E. Jones
* ''Penstemon albomarginatus'' M.E. Jones – White margin penstemon, whitemargin beardtongue, whitemargin penstemon
* ''Penstemon alluviorum'' Pennell – lowland beardtongue
* ''Penstemon ambiguus'' Torr. – bush penstemon, gilia beardtongue, gilia penstemon
* '' Penstemon ammophilus'' N. Holmgren & L. Shultz – sandloving penstemon
* '' Penstemon anguineus'' Eastw. – Siskiyou beardtongue
* ''Penstemon angustifolius'' Nutt. ex Pursh – broad-beard beardtongue, broadbeard beardtongue, narrowleaf penstemon
* '' Penstemon arenarius'' Greene – Nevada sanddune beardtongue, sanddune penstemon
* '' Penstemon arenicola'' A. Nels. – sand penstemon
* '' Penstemon aridus'' Rydb. – stiffleaf penstemon
* '' Penstemon arkansanus'' Pennell – Arkansas beardtongue
* ''Penstemon attenuatus