Peninsular Chuckwalla
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''Sauromalus ater'', also known as the common chuckwalla, is a species of
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
in the family
Iguanidae The Iguanidae is a family of lizards composed of the iguanas, chuckwallas, and their prehistoric relatives, including the widespread green iguana. Taxonomy Iguanidae is thought to be the sister group to the collared lizards (family Crotaph ...
. It inhabits the Sonoran and
Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...
s of the
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...
and northwestern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
. Its range extends from eastern
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
,
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
, and
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
south to
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
and
Sonora Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is d ...
.


Taxonomy

The common name "chuckwalla" (or chuckawalla) is derived from the
Shoshone The Shoshone or Shoshoni ( or ) are a Native American tribe with four large cultural/linguistic divisions: * Eastern Shoshone: Wyoming * Northern Shoshone: southern Idaho * Western Shoshone: Nevada, northern Utah * Goshute: western Utah, easter ...
word ''tcaxxwal'', or ''caxwal'', the form used by the
Cahuilla The Cahuilla , also known as ʔívil̃uqaletem or Ivilyuqaletem, are a Native American people of the various tribes of the Cahuilla Nation, living in the inland areas of southern California. Its
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name, ''Sauromalus'', is said to be a combination of two
ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
words: ''sauros'' meaning "lizard" and ''omalus'' meaning "flat". The proper ancient Greek word for "flat" is however ''homalos'' (ὁμαλός) or ''homalēs'' (ὁμαλής).Liddell, H.G. & Scott, R. (1940). ''A Greek-English Lexicon. revised and augmented throughout by Sir Henry Stuart Jones. with the assistance of. Roderick McKenzie.'' Oxford: Clarendon Press. Its
specific Specific may refer to: * Specificity (disambiguation) * Specific, a cure or therapy for a specific illness Law * Specific deterrence, focussed on an individual * Specific finding, intermediate verdict used by a jury in determining the fina ...
name is ''ater'',
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for "black" or "dark" Its original
epithet An epithet (, ), also byname, is a descriptive term (word or phrase) known for accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, di ...
was ''Sauromalus obesus''; although that name is no longer officially recognized, it is still very common in the literature and it remains in many standard natural history references for North America. In 1998, Bradford D. Hollingsworth examined variations in ''Sauromalus'' and concluded that only five species should be recognized. He regarded ''S. obesus'' as conspecific with ''S. ater'', and he used ''S. ater'', which has priority, as the specific name of the combined taxon. No subspecies of ''S. ater'' are currently recognized. Based primarily on the extensive use of the name ''S. obesus'', a petition to give that name precedence over that of ''S. ater'' was submitted to the
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the I ...
. However, this reasoning was dubious and the priority of ''S. ater'' was maintained. In 2004, ICZN ruled that the name ''Sauromalus ater'' was first described by zoologist
Auguste Duméril Auguste Henri André Duméril (30 November 1812 – 12 November 1870) was a French zoologist. His father, André Marie Constant Duméril (1774-1860), was also a zoologist. In 1869 he was elected as a member of the Académie des sciences. Duméril ...
in 1856, thus had precedence over the name ''Sauromalus obesus'' which was not named until 1858 by Baird.


Description

The common chuckwalla is a large, flat-bodied lizard with a large, rounded belly, and a wide-based, blunt-tipped tail. Reaching a total length of 20 in and a weight of . Small scales cover its body, with larger scales protecting the ear openings. The coloration of these lizards varies by location and between juveniles and adults, as well as between males and females. In adult males, the head, shoulder, and pelvic regions are black, while the midbody is light tan speckled with brown. Adult females are brownish in color with a scattering of dark red spots. Young chuckwallas have four or five broad bands across their bodies, and three or four on the tail which are lost in adulthood by males, but retained somewhat by females.


Behavior

Harmless to humans, these lizards are known to run from potential threats. When disturbed, the chuckwalla enters crevices between rocks and inflates its lungs using a gular pump.Deban, S.M., J.C. O’Reilly, and T.C. Theimer 1994. Mechanism of defensive inflation in the chuckwalla, Sauromalus obesus. Journal of Experimental Zoology 270: 451-459. Lung inflation distends its body and wedges the lizard tightly in place.Stebbins, Robert C.,(2003) ''A Field Guide to Western Reptiles and Amphibians'', 3rd Edition. Houghton Mifflin Company, Males are seasonally and conditionally territorial; an abundance of resources tends to create a hierarchy based on size, with one large male dominating the area's smaller males. Chuckwallas use a combination of color and physical displays, namely "push ups", head-hobbing, and gaping of the mouth to
communicate Communication (from la, communicare, meaning "to share" or "to be in relation with") is usually defined as the transmission of information. The term may also refer to the message communicated through such transmissions or the field of inquir ...
and defend their territory. Chuckwallas are diurnal animals, and as they are
ectothermic An ectotherm (from the Greek () "outside" and () "heat") is an organism in which internal physiological sources of heat are of relatively small or of quite negligible importance in controlling body temperature.Davenport, John. Animal Life a ...
, spend much of their mornings and cooler days basking. These lizards are well adapted to desert conditions; they are active at temperatures up to 102 °F (39 °C). Chuckwallas
brumate Dormancy is a period in an organism's life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolic activity and therefore helps an organism to conserve energy. Dormancy tends to be clos ...
during cooler months and emerge in February. Mating occurs from April to July, with five to 16 eggs laid between June and August. The eggs hatch in late September.


Diet

Primarily
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, the chuckwalla eats
creosote bush ''Larrea tridentata'', called creosote bush and greasewood as a plant, chaparral as a medicinal herb, and ''gobernadora'' (Spanish language, Spanish for "governess") in Mexico, due to its ability to secure more water by inhibiting the growth of n ...
flowers, leaves, fruit, and occasionally insects.


Gallery

File:Sauromalus_ater_1.jpg File: Juvenile common chuckwalla.jpg File:Sauromalus ater 2.jpg File:Sauromalus ater (3).JPG


References


External links


Taxonomic controversyNature Serve
{{Taxonbar, from=Q656553
ater Ater (Hebrew אֲתַר) is an Old Testament male name. #A descendant of Hezekiah, who returned from Babylon ; #An Israelite, who subscribed to Nehemiah’s covenant #A porter ; {{bibleverse, , Nehemiah, 7:45 Set index articles on Hebrew Bible ...
Reptiles described in 1856 Taxa named by Auguste Duméril Reptiles of Mexico Reptiles of the United States Fauna of the Sonoran Desert Fauna of the Mojave Desert Fauna of the Colorado Desert Fauna of the Southwestern United States