Peltophryne Empusa
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The Cuban small-eared toad (''Peltophryne empusa''), also known as the Cuban toad or Cope's Caribbean toad, is a species of
toad Toad is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands. A distinction between frogs and toads is not made in scientif ...
in the family
Bufonidae {{Cat main, Toad This category contains both species commonly called toads, and the true toads from the family Bufonidae {{Cat main, Toad This category contains both species commonly called toads, and the true toads from the family Bufonidae {{Cat m ...
that is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to Cuba including
Isla de Juventud Isla de la Juventud (; en, Isle of Youth) is the second-largest Cuban island (after Cuba's mainland) and the seventh-largest island in the West Indies (after mainland Cuba itself, Hispaniola, Jamaica, Puerto Rico, Trinidad, and Andros Islan ...
.


Description

Species description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have be ...
of ''Peltophryne empusa'' was published by
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontologist, comparative anatomist, herpetologist, and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested ...
in 1862 as an addendum to his work entitled "Notes upon some reptiles of the Old World" (
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s and amphibians were not necessarily considered very distinct at that time):
Supraorbital ridge The brow ridge, or supraorbital ridge known as superciliary arch in medicine, is a bony ridge located above the eye sockets of all primates. In humans, the eyebrows are located on their lower margin. Structure The brow ridge is a nodule or crest ...
s very prominent, not crenate, presenting a posterior process. Postorbital and supra-tympanic processes prominent, obtuse; preorbital straight, more acute. Canthus rostrales acute, converging so as to produce a very acute angle; their profile very declive, that of the muzzle more so, but not perpendicular. Maxillary region oblique from a front view; the labial border forming a prominent rim, which is thickened and everted posteriorly. Two occipital knobs on each side. Tympanum small, one-fourth or one-third the length of the palpebral border in diameter.
Paratoid gland The parotoid gland (alternatively, paratoid gland) is an external skin gland on the back, neck, and shoulder of toads and some frogs and salamanders. It can secrete a number of milky alkaloid substances (depending on the species) known collective ...
small, rounded, lateral, studded with warts; the dorsal region is similarly studded, most abundantly anteriorly. Sides, extremities and gular region covered with smaller warts; belly areolate. One large oval flat metacarpal tubercle; a large one at the base of the interior digit. Two metatarsal tubercles; the interior most elongate and acute, blackish brown. A short, thickened, internal tarsal fold. Toes half-webbed, palm slightly rugose. Length from end of muzzle to tympanum 11
lines Line most often refers to: * Line (geometry), object with zero thickness and curvature that stretches to infinity * Telephone line, a single-user circuit on a telephone communication system Line, lines, The Line, or LINE may also refer to: Arts ...
; of antebrachium and hand, 14.5 lines;
axilla The axilla (also, armpit, underarm or oxter) is the area on the human body directly under the shoulder joint. It includes the axillary space, an anatomical space within the shoulder girdle between the arm and the thoracic cage, bounded superior ...
to vent, 2 inches; vent to end of fourth toe, 3 inches 1 line. The head is brown; color elsewhere brownish yellow; on the nape and sides marbled with deep brown, somewhat oblique-longitudinally on the latter region. Limbs cross-banded with brown.


Distribution and habitat

''Peltophryne empusa'' has a wide but patchy distribution in
xeric Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (ancient Greek xērós, “dry") shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this h ...
and mesic lowland forests and
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
s of Cuba and the Isla de Juventud to
asl American Sign Language (ASL) is a natural language that serves as the predominant sign language of Deaf communities in the United States of America and most of Anglophone Canada. ASL is a complete and organized visual language that is express ...
. However, it burrows underground and is rarely seen except during the breeding season when it is abundant. It is an explosive breeder; males call from flooded ditches and large temporary pools of rainwater. Eggs are laid in still water.


Conservation

''Peltophryne empusa'' is assessed as a
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
because its distribution area is less than 2,000 km² and severely fragmented, and it is affected by habitat loss and degradation caused by agriculture, pollution (pesticides), and the invasive
legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock f ...
''
Dichrostachys cinerea ''Dichrostachys cinerea'', known as sicklebush, Bell mimosa, Chinese lantern tree or Kalahari Christmas tree (South Africa), is a legume of the genus ''Dichrostachys'' in the family Fabaceae. Other common names include omubambanjobe (Tooro Ugan ...
''. However, it occurs in several protected areas.


References

{{Taxonbar, from=Q2236367 Peltophryne Amphibians described in 1862 Endemic fauna of Cuba Amphibians of Cuba Taxa named by Edward Drinker Cope Taxonomy articles created by Polbot