Pelargonium Luridum
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''Pelargonium luridum'', locally called variable stork's bill, is a medium high, tuberous
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
perennial
geophyte A storage organ is a part of a plant specifically modified for storage of energy (generally in the form of carbohydrates) or water. Storage organs often grow underground, where they are better protected from attack by herbivores. Plants that have ...
, belonging to the Stork's bill family, with white to pink, slightly mirror symmetrical flowers in umbels on long unbranched stalks directly from the ground rosette that consists of few initially ovate, later pinnately incised or linear leaves, with blunt teeth around the margin. The variable stork's bill naturally occurs from South Africa to Angola, southern Congo and Tanzania.


Description

''Pelargonium luridum'' is a very variable species. It is an erect, perennial, herbaceous plant with up to long inflorescence stalks, with a woody tuber and with all leaves growing directly from the hart of the plant at ground level. The plants are covered in a layer of soft, about long hairs, and shorter hairs that are pressed to the surface of the plant. It also has glands. Most of the leaves appear after flowering. The stipules on both sides next to the base of the leaf stalks are linear or long triangular in shape and long. These are persistent and turn brown with age. The leaf stalks are usually , occasionally up to long. The leaf blade is oval in outline, usually long and wide, wedge- to hart-shaped at its foot, variably incised, form shallowly lobed to multiply dissected, the later leaves often more complex. The segments at the margins of the leaf-outline, can be thread-thin to oval, entire of with teeth around the margin, and the tip pointy of rounded. The upper leaf surface may be sofly hairy or almost without, except for the margins and veins. Each plant may carry one to three inflorescences, consisting of a long peduncle with few to many small, pointy, simple membraneous bracts long and 1½–4 mm (0.06–0.16 in) wide, topped by a simple umbel of usually between five and thirty, sometimes even fifty flowers. At the top of the peduncle is a whorl of small tapered bracts. The lower part of the stalk of the individual flower is mostly long. The sepals are deflected when the flower has opened, lanceolate to narrowly oval, 8½–15 mm (0.33–0.6 in) long and 1½–5 mm (0.06–0.20 in) wide, and are covered in dense felty hairs and glands. The flowers have a spur, that has merged with the upper part of the flower stalk of long. There are five
petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...
s present, which are at first spreading but later somewhat reflexed, and are long inverted egg-shaped or lanceolate with the widest point towards the tip, usually long and wide. Their color varies from white to pale yellow, with a pink venation or entirely purple. The three anterior petals have a narrow basal half (a so-called claw) and a wider top half (or plate). The ten stamens are fused at their foot for 1¾–3½ mm (0.07–0.14 in). Usually seven of these stamens carry long, 1¼–1¾ mm (0.05–0.07 in) wide
anthers The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
, and their filaments are long, the usually three infertile
staminode In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co. ...
s are 3–5½ mm (0.12–0.22 in) long. The ovary is softly hairy, with a short style long, topped by five stigmas long. It develops into a fruit that grows to , exceptionally 6½ cm long. When ripe each of the five seeds comes of with a long and 1½–3 mm (0.06–0.12 in) wide part of the fruit called a
mericarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the m ...
. Individual seeds within the mericarp are long and wide. The pale brown seed coat has a very fine netted structure.


Taxonomy

Henry Cranke Andrews was the first to describe this species in 1813 as ''Geranium luridum''. In 1820, Robert Sweet described a non-flowering plant that was grown in England by a mrs. Colville, from a rootstock sent from South Africa, calling it ''Pelargonium huraefolium''. After the plant had started flowering later, Sweet realised his species was in fact identical to that of Andrews, but remained of the opinion that it belonged to ''Pelargonium'', and so created the
new combination ''Combinatio nova'', abbreviated ''comb. nov.'' (sometimes ''n. comb.''), is Latin for "new combination". It is used in taxonomic biology literature when a new name is introduced based on a pre-existing name. The term should not to be confused wi ...
''Pelargonium luridum'' in 1826. ''Polyactium aconitiphyllum'' was described based on a specimen from South Africa in 1835 by
Christian Friedrich Ecklon Christian Friedrich Ecklon (17 December 1795 – 1 December 1868) was a Denmark, Danish botany, botanical collector and apothecary. Ecklon is especially known for being an avid collector and researcher of plants in South Africa. Biography Ecklon ...
and
Karl Ludwig Philipp Zeyher Karl Ludwig Philipp Zeyher (2 August 1799 Dillenburg, Hessen, Germany – 13 December 1858 Cape Town), was a botanical and insect collector who collected extensively in South Africa. He was the author, with Christian Friedrich Ecklon, of ''Enumer ...
. Ernst Gottlieb von Steudel moved it in 1842 to ''Pelargonium'', creating the new combination ''Pelargonium aconitiphyllum''. Because the species is very variable, several later authors raised different morphs to their own species. Ernst Heinrich Friedrich Meyer in 1843 named ''P. polymorphum'', but failed to adequately describe it. In 1860 William Henry Harvey distinguished ''P. flabellifolium'', and Harvey in cooperation with Otto Wilhelm Sonder defined ''Pelargonium zeyheri'' in the same year.
Ignaz von Szyszyłowicz Ignaz von Szyszyłowicz (30 July 1857 – 17 February 1910) also known as Ignacy Szyszyłowicz was a Polish botanist born in Granica (Sosnowiec). He contributed Part III.6 ''Caryocaraceae, Marcgraviaceae, Theaceae, Strasburgeriaceae'' to Engler ...
distinguished two other forms, naming them ''P. rehmannii'' and ''P. angulosum'' respectively in the year 1888.
Otto Kuntze Carl Ernst Otto Kuntze (23 June 1843 – 27 January 1907) was a German botanist. Biography Otto Kuntze was born in Leipzig. An apothecary in his early career, he published an essay entitled ''Pocket Fauna of Leipzig''. Between 1863 and 1866 he ...
moved ''flabellifolium'', ''rehmannii'' and ''zeyheri'' to a new, more narrowly defined genus ''Geraniospermum'', in 1891. In 1902, Adolf Engler describes ''P. heckmannianum''. All of these names are now considered
synonymous A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
to ''P. luridum''. ''P. luridum'' is the only species assigned to the subsection Magnistipulacea, which itself is part of the large section Polyactium.


Etymology

The species name ''luridum'' is from Latin and means pale, sallow or smoky yellow.


Distribution and habitat

''Pelargonium luridum'' occurs naturally in Angola, Eswatini, Lesotho, Malawi, South-Africa (Cape Province, Natal, Orange Free State, Transvaal), Tanzania, Zambia, Zimbabwe. It can be found in grassland and open bush, is said to prefer locations subject to regular fires, and damp ground such as by streams.


References

{{Taxonbar, from=Q7213346 Plants described in 1813 luridum Flora of East Tropical Africa Flora of South Tropical Africa Flora of Southern Africa Taxa named by Henry Cranke Andrews