Parcoblatta Americana
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''Parcoblatta americana'', the western wood cockroach, is a species of wood
cockroach Cockroaches (or roaches) are a Paraphyly, paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are we ...
that occurs in Mexico and the western United States.


Description

The species has significant size and color variation, thought to be due to environmental conditions, particularly the effect of aridity. The male of the species has normal, fully developed
tegmina A tegmen (plural: ''tegmina'') designates the modified leathery front wing on an insect particularly in the orders Dermaptera ( earwigs), Orthoptera (grasshoppers, crickets and similar families), Mantodea (praying mantis), Phasmatodea (stick an ...
and wings, and a tuft of hairs on the back of its middle abdomen. The general coloration in the male can vary from shining dark brown or blackish to a pale yellowish to slightly reddish tan. The tegmina are translucent in all colorations, and the
ocelli A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-l ...
(simple eye spots) are a yellowish tan. Females of the species have very greatly reduced tegmina, represented by lobe-like lateral pads, and no hind wings. The female general coloration ranges from a shining reddish orange or brown, slightly darker on the back of the abdomen, to a shining dark brown, or shining black with blackish brown underparts. The ocelli are tannish in all colorations.
Morgan Hebard Morgan Hebard (February 23, 1887 – December 28, 1946) was an American entomologist who specialized in orthoptera, and assembled a collection of over 250,000 specimens. Early life and education Morgan Hebard was born on February 23, 1887, in Cl ...
's 1917 description included measurement ranges based on 14 male specimens and 7 female specimens: :


Distinguishing characteristics

The male ''P. americana'' has a single specialized area on its abdomen, on the back of its median segment, in the form of a tuft of agglutinated hairs. ''Parcoblatta zebra'' is the only other male of the genus ''Parcoblatta'' that shares this characteristic, but on ''P. americana'' the tuft is small and quadrate (squarish), with the rest of the segment showing little specialization, while on ''P. zebra'' the tuft is much broader, with the rest of the segment showing further specialization. The male ''Parcoblatta virginica'' also has a single specialized area on its abdomen, but it has minute hairs scattered over a large specialized area on the back of its median segment, rather than a tuft of agglutinated hairs. The female ''Parcoblatta bolliana'' is the only other female of the genus ''Parcoblatta'' that has such significantly reduced tegmina and absence of hind wings. The female ''P. bolliana'' is distinguished from ''P. americana'' by its compact form, space between its compound eyes less than the space between its antennal sockets, its tegmina having few traces of
venation Venation may refer to: * Venation (botany), the arrangement of veins in leaves * Wing venation, the arrangement of veins in insect wings See also * * Vernation Vernation (from ''vernal'' meaning ''spring'', since that is when leaves spring fort ...
, and its supra-anal plate weakly produced with its side edges converging to a broadly rounded apex, while the ''P. americana'' having a normal, less compact form, space between its compound eyes noticeably wider than the space between its antennal sockets, its tegmina have visible venation, its supra-anal plate is normally produced with its side edges converging to a sharply rounded apex.


Distribution and habitat

The species is known in
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
and the western United States, including Oregon, California, Nevada, and Arizona. It has also been recorded in a home in
Klickitat County, Washington Klickitat County is a county located in the U.S. state of Washington. As of the 2020 census, the population was 22,735. The county seat and largest city is Goldendale. The county is named after the Klickitat tribe. History Klickitat Count ...
. Both genders of ''P. americana'' have been found in the nests of the ''Veromessor chicoensis'' (a
harvester ant Harvester ant, also known as harvesting ant, is a common name for any of the species or genera of ants that collect seeds (called seed predation), or mushrooms as in the case of '' Euprenolepis procera'', which are stored in the nest in commu ...
, also classified as ''Messor chicoensis''), where they reside during the day, and emerge at night to feed on the debris pile around the nest.
Nymphs A nymph ( grc, νύμφη, nýmphē, el, script=Latn, nímfi, label=Modern Greek; , ) in ancient Greek folklore is a minor female nature deity. Different from Greek goddesses, nymphs are generally regarded as personifications of nature, are ...
of ''P. americana'' were taken from houses of
pack rat A pack rat or packrat, also called a woodrat or trade rat, are any species in the North and Central American rodent genus ''Neotoma''. Pack rats have a rat-like appearance, with long tails, large ears, and large, black eyes. Pack rats are notice ...
s (genus ''Neotoma'') in
Orange County, California Orange County is located in the Los Angeles metropolitan area in Southern California. As of the 2020 census, the population was 3,186,989, making it the third-most-populous county in California, the sixth-most-populous in the United States, a ...
. The species has been observed feeding on an apple that was six feet off the ground. A study area in California's
Santa Monica Mountains The Santa Monica Mountains is a coastal mountain range in Southern California, next to the Pacific Ocean. It is part of the Transverse Ranges. Because of its proximity to densely populated regions, it is one of the most visited natural areas in ...
found the species to be fairly common, but localized to the area. The study found them occurring in chaparral, grassland, oak woodland, and coastal sage (''Artemisia californica''), and occurring as adults in early to mid summer. Females and nymphs lived on the ground, and were found under rocks, in rotting wood, in ant nests. Adult males lived separately from females and nymphs; they were attracted to light, and would sometimes perch on chaparral plants at night.


References


External links


Drawings
from a 1917 article by Morgan Hebard. Plate III includes dorsal views of male and female specimens of ''P. americana'' (figures 7 and 9), and a view of the underside of the male subgenital plate (figure 8). Key to drawings on page 276.
Black and white photographs
of top view of ''P. americana'' male and female specimens, from ''Smithsonian Miscellaneous Collections''. {{Taxonbar, from=Q10620165 Cockroaches Insects of North America Insects of Mexico Insects of the United States Insects described in 1900