Paranibea
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The half-mourning croaker (''Paranibea semiluctuosa'') is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
belonging to the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Sciaenidae Sciaenidae are a family of fish in the order Acanthuriformes. They are commonly called drums or croakers in reference to the repetitive throbbing or drumming sounds they make. The family consists of about 286 to 298 species in about 66 to 70 gen ...
, the drums and croakers. It is the only species in the
monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
''Paranibea''. This fish is found in the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region.


Taxonomy

The half-mourning croaker was first formally described as ''Corvina semiluctuosa'' in 1830 by the French
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...
with
Malabar Malabar may refer to the following: People * Malabars, people originating from the Malabar region of India * Malbars or Malabars, people of Tamil origin in Réunion Places * Malabar Coast, or Malabar, a region of the southwestern shoreline o ...
given as the type locality. In 1977
Ethelwynn Trewavas Ethelwynn Trewavas (5 November 1900 – 16 August 1993) was an ichthyologist at the British Museum of Natural History. She was known for her work on the families Cichlidae and Sciaenidae. She worked with Charles Tate Regan, another ichthyo ...
classified ''C. semiluctuosa'' in a new monospecific genus ''Paranibea''. This taxon is included in the subfamily Otolithinae by some workers, but the 5th edition of ''
Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...
'' does not recognise subfamilies within the Sciaenidae, which it places in the
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
Acanthuriformes Acanthuriformes is an order of ray-finned fishes, part of the Percomorpha clade. Some authorities place the fishes in the order within the Acanthuriformes in the suborders Acanthuroidea and Percoidea of the order Perciformes. Classification The ...
.


Etymology

The half-mourning croaker was considered to be classified within '' Nibea'' and the genus name, ''Paranibea'', means "like" or "close to" ''Nibea''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''semiluctuosa'' translates as "half mourning" and is presumed to refer to the dark stripes running along the scales over most of the body, except for a narrow area on the belly.


Description

The half-mourning croaker has a deep body, which is over 3 times the
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
, with a rounded dorsal profile and a slightly inferior mouth, which has thick, warty lips. The teeth in the lower jaw are all the same size. The
swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled Organ (anatomy), organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their curren ...
is shaped like a carrot and has branched appendages along the entirety of each side, the pair at the front go through the
transverse septum The septum transversum is a thick mass of cranial mesenchyme, formed in the embryo, that gives rise to parts of the thoracic diaphragm and the ventral mesentery of the foregut in the developed human being and other mammals. Origins The septum tr ...
. The
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
has a shallow notch with 9 or 10 spines before the notch and 1 spine and between 27 and 31 soft rays to its rear. The
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
is supported by 2 spines and 7 or 8 soft rays. The colour is dark with numerous, slender, sinuous black stripes and with the anal and
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
s being darker than the other fins. The maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
for this species is , although is more typical.


Distribution and habitat

The half-mourning croaker is found in the Indo-West Pacific where it has it is found in Iran and Pakistan east to northern Myanmar, it has been recorded in
Peninsular Malaysia Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
but records from farther east require to be verified. It is found in coastal waters over mud substrates, although in Bangladesh it has been recorded on sandy substrates.


Fisheries

The half-mourning croaker is commonly landed by commercial fisheries, especially in western India, and is also taken as
bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
. The fish landed are sold fresh or are preserved by
drying Drying is a mass transfer process consisting of the removal of water or another solvent by evaporation from a solid, semi-solid or liquid. This process is often used as a final production step before selling or packaging products. To be considered ...
or salting.


References

{{Taxonbar, from=Q1989924, from2=Q2496112 Sciaenidae Taxa named by Ethelwynn Trewavas Taxa named by Georges Cuvier Fish described in 1830