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The stone sculpin (''Paracottus knerii'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of cottoid fish
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
, where it is found in
Lake Baikal Lake Baikal (, russian: Oзеро Байкал, Ozero Baykal ); mn, Байгал нуур, Baigal nuur) is a rift lake in Russia. It is situated in southern Siberia, between the federal subjects of Irkutsk Oblast to the northwest and the Repu ...
and surrounding tributaries as well as the Gramninskie Lakes, Lake Verkhnaya Agata and the
Enisei River The Yenisey (russian: Енисе́й, ''Yeniséy''; mn, Горлог мөрөн, ''Gorlog mörön''; Buryat: Горлог мүрэн, ''Gorlog müren''; Tuvan: Улуг-Хем, ''Uluğ-Hem''; Khakas: Ким суғ, ''Kim suğ''; Ket: Ӄук ...
and various lakes in
Tuva Tuva (; russian: Тува́) or Tyva ( tyv, Тыва), officially the Republic of Tuva (russian: Респу́блика Тыва́, r=Respublika Tyva, p=rʲɪˈspublʲɪkə tɨˈva; tyv, Тыва Республика, translit=Tyva Respublika ...
. This species is the only recognized member of its
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
. It is often eaten by the
Baikal seal The Baikal seal, Lake Baikal seal or (''Pusa sibirica''), is a species of earless seal endemic to Lake Baikal in Siberia, Russia. Like the Caspian seal, it is related to the Arctic ringed seal. The Baikal seal is one of the smallest true seals a ...
, especially in the autumn.


Taxonomy

The stone sculpin was first formally described as ''Cottus knerii'' in 1874 by the Polish
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Benedykt Dybowski Benedykt Tadeusz Dybowski (12 May 183331 January 1930) was a Polish naturalist and physician. Life Benedykt Dybowski was born in Adamaryni, within the Minsk Governorate of the Russian Empire to Polish nobility. He was the brother of naturalist ...
with its type locality given as Lake Baikal and the
Angara The Angara ( Buryat and mn, Ангар, ''Angar'',  "Cleft"; russian: Ангара́, ''Angará'') is a major river in Siberia, which traces a course through Russia's Irkutsk Oblast and Krasnoyarsk Krai. It drains out of Lake Baikal and is ...
, Irkut and
Selenga The Selenga or Selenge ( ; bua, Сэлэнгэ гол / Сэлэнгэ мүрэн, translit=Selenge gol / Selenge müren; russian: Селенга́, ) is a major river in Mongolia and Buryatia, Russia. Originating from its headwater tributaries, ...
rivers in
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
. In 1949
Dmitrii Nikolaevich Taliev Dmitrii Nikolaevich Taliev (28 May 1908 – 2 July 1952) was a Soviet Russian ichthyologist and limnologist, notable for his work on the Lake Baikal. Notable Dates (includes list of publications and short biography) * 1930 - Senior Assistant, P ...
classified the stone sculpin in the
monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
''Porocottus''. The genus was classified in the family Cottocomephoridae, however, the 5th edition of ''
Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...
'' classifies the genus within the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Cottinae Cottinae is a subfamily of ray-finned fishes belonging to the family Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. ...
of the family Cottidae, studies having found that the genera formerly included in the Cottocomephoridae were a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
arising from the genus '' Cottus''.


Appearance

Maximum total length of the stone sculpin is , but most are . Its colour varies from
olive The olive, botanical name ''Olea europaea'', meaning 'European olive' in Latin, is a species of small tree or shrub in the family Oleaceae, found traditionally in the Mediterranean Basin. When in shrub form, it is known as ''Olea europaea'' ...
to brown or grey, and it has several dark spots and bands.


Habitat

The stone sculpin is most common in the southern part of Lake Baikal and prefers areas with a stony bottom, but may also occur over sand. It is typically found from shallow water to a depth of , but can occur to .


Behavior

The stone sculpin breeds in the late spring and summer in water that is , and up to deep. The female lays 200–700 eggs in a crevice under a stone, which are guarded by the male and hatch after about one month. This sculpin feeds on small animals such as
copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...
s and
amphipod Amphipoda is an order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods range in size from and are mostly detritivores or scavengers. There are more than 9,900 amphipod species so far descr ...
s.


References

{{Taxonbar, from=Q6407887 Cottinae Monotypic fish genera Fish of Lake Baikal Fish described in 1874