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''Panthera zdanskyi'', also known as the Longdan tiger, is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
pantherine
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
that is seen as a close relative of the modern
tiger The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on u ...
.
Fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
were excavated in northwestern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
's
Gansu province Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibeta ...
.


Etymology

''Panthera zdanskyi'' was first named by Ji H. Mazák, Per Christiansen and Andrew C. Kitchener in
2011 File:2011 Events Collage.png, From top left, clockwise: a protester partaking in Occupy Wall Street heralds the beginning of the Occupy movement; protests against Libyan dictator Muammar Gaddafi, who was killed that October; a young man celebrate ...
. The specific name honors the Austrian paleontologist Otto A. Zdansky for his contributions to the understanding of
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...
Chinese fossil carnivorans.


Description

The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
of ''Panthera zdanskyi'' consists of a nearly complete
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
and
mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
stored at the Babiarz Institute of Paleontological Studies. It is the oldest known complete pantherine skull ever found, and indicates that the animal was about the size of a
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
. The
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Of ...
comprises a
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ships * Ros ...
,
premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has b ...
and
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
and much of the dentition, originally referred to ''Panthera palaeosinensis''. It was collected in 2004 in the east slope of Longdan, south of
Dongxiang Autonomous County Dongxiang Autonomous County (; Santa: Dunxianzu Zizhixien) is an autonomous county in the Linxia Hui Autonomous Prefecture, province of Gansu of the People's Republic of China. It was established as a Dongxiang ethnic autonomous area in 1950. Hist ...
in the Lower Pleistocene ''Equus'' layer, dating to the
Gelasian The Gelasian is an age (geology), age in the international geologic timescale or a stage (stratigraphy), stage in chronostratigraphy, being the earliest or lowest subdivision of the Quaternary Period/System and Pleistocene Epoch/Series. It spans ...
stage of the earliest
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
, about 2.55–2.16
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
.


Phylogeny

The
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
below follows Mazák, Christiansen and Kitchener (2011).


See also

*''
Panthera blytheae ''Panthera blytheae'' is an extinct species of the genus '' Panthera'' that is thought to have existed during the late Messinian to early Zanclean ages approximately 5.95–4.1 million years ago. It is currently the oldest known ''Panthera'' sp ...
'' *''
Panthera gombaszoegensis ''Panthera gombaszoegensis'', also known as the European jaguar, is a ''Panthera'' species that lived from about 2.0 to 0.35 million years ago in Europe. The first fossils were excavated in 1938 in Gombasek, Slovakia. More of its fossil remai ...
'' *'' Panthera shawi'' *'' Panthera sondaica'' *''
Panthera youngi ''Panthera youngi'' is a fossil cat species that was described in 1934; fossil remains of this cat were excavated in a ''Sinanthropus'' formation in Choukoutien, northeastern China. Upper and lower jaws excavated in Japan's Yamaguchi Prefecture ...
''


References


External links

zdanskyi Fossil taxa described in 2011 zdanskyi Pleistocene carnivorans Pleistocene mammals of Asia {{felidae-stub