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The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear (or simply the panda), is a
bear Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Nor ...
species
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. It is characterised by its bold black-and-white
coat A coat typically is an outer garment for the upper body as worn by either gender for warmth or fashion. Coats typically have long sleeves and are open down the front and closing by means of buttons, zippers, hook-and-loop fasteners, toggles, a ...
and rotund body. The name "giant panda" is sometimes used to distinguish it from the
red panda The red panda (''Ailurus fulgens''), also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle ...
, a neighboring musteloid. Though it belongs to the order
Carnivora Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...
, the giant panda is a
folivore In zoology, a folivore is a herbivore that specializes in eating leaves. Mature leaves contain a high proportion of hard-to-digest cellulose, less energy than other types of foods, and often toxic compounds.Jones, S., Martin, R., & Pilbeam, D. (1 ...
, with
bamboo Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
shoots and leaves making up more than 99% of its diet. Giant pandas in the wild occasionally eat other grasses, wild tubers, or even meat in the form of birds, rodents, or carrion. In captivity, they may receive honey, eggs, fish, yams,
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
leaves, oranges, or bananas along with specially prepared food. The giant panda lives in a few mountain ranges in central China, mainly in
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
, and also in neighbouring
Shaanxi Shaanxi (alternatively Shensi, see #Name, § Name) is a landlocked Provinces of China, province of China. Officially part of Northwest China, it borders the province-level divisions of Shanxi (NE, E), Henan (E), Hubei (SE), Chongqing (S), Sichu ...
and
Gansu Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibet ...
. As a result of farming,
deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated d ...
, and other development, the giant panda has been driven out of the lowland areas where it once lived, and it is a conservation-reliant
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
. A 2007 report showed 239 pandas living in captivity inside China and another 27 outside the country. By December 2014, 49 giant pandas lived in captivity outside China, living in 18 zoos in 13 countries. Wild population estimates vary; one estimate shows that there are about 1,590 individuals living in the wild, while a 2006 study via
DNA analysis Genetic testing, also known as DNA testing, is used to identify changes in DNA sequence or chromosome structure. Genetic testing can also include measuring the results of genetic changes, such as RNA analysis as an output of gene expression, or ...
estimated that this figure could be as high as 2,000 to 3,000. Some reports also show that the number of giant pandas in the wild is on the rise. By March 2015, the wild giant panda population had increased to 1,864 individuals. In 2016, it was reclassified on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
from "endangered" to "vulnerable", affirming decade-long efforts to save the panda. In July 2021, Chinese authorities also reclassified the giant panda as vulnerable. The giant panda has often served as China's
national symbol A national symbol is a symbol of any entity considering and manifesting itself to the world as a national community: the sovereign states but also nations and countries in a state of colonial or other dependence, federal integration, or even an ...
, appeared on
Chinese Gold Panda The Chinese Gold Panda () is a series of gold bullion coins issued by the People's Republic of China. The Official Mint of the People's Republic of China introduced the panda gold bullion coins in 1982. The panda design changes every year (with a ...
coins since 1982 and as one of the five
Fuwa The Fuwa (; literally "good-luck dolls", also known as "Friendlies") were the mascots of the 2008 Summer Olympics in Beijing. The designs were created by Han Meilin, a famous Chinese artist. The designs were publicly announced by the Nationa ...
mascots of the
2008 Summer Olympics The 2008 Summer Olympics (), officially the Games of the XXIX Olympiad () and also known as Beijing 2008 (), were an international multisport event held from 8 to 24 August 2008, in Beijing, China. A total of 10,942 athletes from 204 Na ...
.


Taxonomy


Classification

For many decades, the precise
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
classification of the giant panda was under debate because it shares characteristics with both bears and
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
s. However, molecular studies indicate the giant panda is a true bear, part of the family Ursidae. These studies show it diverged about from the
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. All living beings are in fact descendants of a unique ancestor commonly referred to as the last universal comm ...
of the
Ursidae Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the Nort ...
; it is the most basal member of this family and equidistant from all other extant bear species. The giant panda has been referred to as a
living fossil A living fossil is an extant taxon that cosmetically resembles related species known only from the fossil record. To be considered a living fossil, the fossil species must be old relative to the time of origin of the extant clade. Living fossi ...
.


Etymology

The word ''panda'' was borrowed into English from French, but no conclusive explanation of the origin of the French word ''panda'' has been found. The closest candidate is the Nepali word ''ponya,'' possibly referring to the adapted wrist bone of the red panda, which is native to Nepal. The Western world originally applied this name to the red panda. In many older sources, the name "panda" or "common panda" refers to the lesser-known
red panda The red panda (''Ailurus fulgens''), also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle ...
, thus necessitating the inclusion of "giant" and "lesser/red" prefixes in front of the names. Even in 2013, the ''
Encyclopædia Britannica The (Latin for "British Encyclopædia") is a general knowledge English-language encyclopaedia. It is published by Encyclopædia Britannica, Inc.; the company has existed since the 18th century, although it has changed ownership various time ...
'' still used "giant panda" or "panda bear" for the bear, and simply "panda" for the
red panda The red panda (''Ailurus fulgens''), also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle ...
, despite the popular usage of the word "panda" to refer to giant pandas. Since the earliest collection of Chinese writings, the Chinese language has given the bear 20 different names, such as '' '' ( ancient Chinese name for giant panda), ''huāxióng'' ( "spotted bear") and ''zhúxióng'' ( "bamboo bear"). The most popular names in China today are ''dàxióngmāo'' ( literally "giant bear cat"), or simply ''xióngmāo'' ( "bear cat"). The name ''xióngmāo'' ( "bear cat") was originally used to describe the
red panda The red panda (''Ailurus fulgens''), also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle ...
(''Ailurus fulgens''), but since the giant panda was thought to be closely related to the red panda, ''dàxióngmāo'' () was named relatively. In
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
, another popular name for panda is the inverted ''dàmāoxióng'' ( "giant cat bear"), though many encyclopediae and dictionaries in Taiwan still use the "bear cat" form as the correct name. Some linguists argue, in this construction, "bear" instead of "cat" is the base noun, making this name more grammatically and logically correct, which may have led to the popular choice despite official writings. This name did not gain its popularity until 1988, when a private zoo in
Tainan Tainan (), officially Tainan City, is a Special municipality (Taiwan), special municipality in southern Taiwan facing the Taiwan Strait on its western coast. Tainan is the oldest city on the island and also commonly known as the "Capital City" ...
painted a
sun bear The sun bear (''Helarctos malayanus'') is a species in the family Ursidae (the only species in the genus ''Helarctos'') occurring in the tropical forests of Southeast Asia. It is the smallest bear, standing nearly at the shoulder and weighing ...
black and white and created the
Tainan fake panda incident The Tainan fake panda incident refers to an incident at a private zoo in Tainan, Taiwan, where a sun bear was painted black and white and falsely presented to visitors as a giant panda. The bear was first unveiled on 24 December 1987, and quickly ...
.


Subspecies

Two subspecies of giant panda have been recognized on the basis of distinct cranial measurements, colour patterns, and
population genetics Population genetics is a subfield of genetics that deals with genetic differences within and between populations, and is a part of evolutionary biology. Studies in this branch of biology examine such phenomena as adaptation, speciation, and pop ...
. * The
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, ''A. m. melanoleuca'', consists of most extant populations of the giant panda. These animals are principally found in
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
and display the typical stark black and white contrasting colours. * The
Qinling panda The Qinling panda (''Ailuropoda melanoleuca qinlingensis'') is a subspecies of the giant panda, discovered in the 1960s but not recognized as a subspecies until 2005. Besides the nominate subspecies, it is the first giant panda subspecies to be r ...
, ''A. m. qinlingensis'', is restricted to the
Qinling Mountains The Qinling () or Qin Mountains, formerly known as the Nanshan ("Southern Mountains"), are a major east–west mountain range in southern Shaanxi Province, China. The mountains mark the divide between the drainage basins of the Yangtze and Yellow ...
in Shaanxi at elevations of . The typical black and white pattern of Sichuan giant pandas is replaced with a light brown and white pattern. The skull of ''A. m. qinlingensis'' is smaller than its relatives, and it has larger molars. A detailed study of the giant panda's genetic history from 2012 confirms that the separation of the Qinlin population occurred about 300,000 years ago, and reveals that the non-Qinlin population further diverged into two groups, named the
Minshan Min Mountains or Minshan () are a mountain range in central China. It runs in the general north-south direction through northern Sichuan (the eastern part of the Ngawa Tibetan and Qiang Autonomous Prefecture and adjacent areas of Mianyang Prefec ...
and the Qionglai-
Daxiangling Daxiangling (), or Daxiang Range is a fairly small mountain range in China's Sichuan Province. It runs in the general western and northwestern direction from the famous sacred Mount Emei, and is part of the western mountainous rim of the Sichuan Bas ...
-
Xiaoxiangling The Xiaoxiang Range () is a mountain range in Sichuan Province, China. It is part of a complicated system of mountains in south-central Sichuan, and runs in the general north-south direction within Liangshan Yi Autonomous Prefecture and the adjace ...
- Liangshan group respectively, about 2,800 years ago.


Phylogeny


Description

Adults measure around long, including a tail of about , and tall at the
shoulder The human shoulder is made up of three bones: the clavicle (collarbone), the scapula (shoulder blade), and the humerus (upper arm bone) as well as associated muscles, ligaments and tendons. The articulations between the bones of the shoulder mak ...
. Males can weigh up to . Females (generally 10–20% smaller than males) can weigh as little as , but can also weigh up to . The average weight for adults is . The giant panda has a body shape typical of bears. It has black fur on its ears, eye patches, limbs and shoulders. The rest of the animal's coat is white. The bear's distinctive coat appears to serve as
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the ...
in both winter and summer environments. The white areas may serve as camouflage in snow, while the black shoulders and legs provide crypsis in shade. Studies in the wild have found that when viewed from a distance, the panda displays
disruptive coloration Disruptive coloration (also known as disruptive camouflage or disruptive patterning) is a form of camouflage that works by breaking up the outlines of an animal, soldier or military vehicle with a strongly contrasting pattern. It is often comb ...
while close up, they rely more on blending in. The black ears may signal aggressive intent, while the eye patches might facilitate them identifying one another. The giant panda's thick, woolly coat keeps it warm in the cool forests of its habitat. The panda's skull shape is typical of
durophagous Durophagy is the eating behavior of animals that consume hard-shelled or exoskeleton bearing organisms, such as corals, shelled mollusks, or crabs. It is mostly used to describe fish, but is also used when describing reptiles, including fossil tu ...
carnivorans. It has evolved from previous ancestors to exhibit larger molars with increased complexity and expanded temporal fossa. A giant panda has a 3D canine teeth bite force of 2603.47 newtons and
bite force quotient Bite force quotient (BFQ) is a numerical value commonly used to represent the bite force of an animal, while also taking factors like the animal's size into account. The BFQ is calculated as the regression of the quotient of an animal's bite f ...
of 292. Another study had a giant panda bite of 1298.9 newtons (BFQ 151.4) at canine teeth and 1815.9 newtons (BFQ 141.8) at carnassial teeth. The giant panda's
paw A paw is the soft foot-like part of a mammal, generally a quadruped, that has claws. Common characteristics The paw is characterised by thin, pigmented, keratinised, hairless epidermis covering subcutaneous collagenous and adipose tissue, w ...
has a "thumb" and five fingers; the "thumb" – actually a modified
sesamoid bone In anatomy, a sesamoid bone () is a bone embedded within a tendon or a muscle. Its name is derived from the Arabic word for ' sesame seed', indicating the small size of most sesamoids. Often, these bones form in response to strain, or can be pres ...
– helps it to hold bamboo while eating.
Stephen Jay Gould Stephen Jay Gould (; September 10, 1941 – May 20, 2002) was an American paleontologist, evolutionary biologist, and historian of science. He was one of the most influential and widely read authors of popular science of his generation. Gould sp ...
discusses this feature in his book of essays on
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
and
biology Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary i ...
, '' The Panda's Thumb''. The giant panda's tail, measuring , is the second-longest in the bear family, behind the
sloth bear The sloth bear (''Melursus ursinus'') is a myrmecophagous bear species native to the Indian subcontinent. It feeds on fruits, ants and termites. It is listed as Vulnerable species, vulnerable on the IUCN Red List, mainly because of habitat loss ...
. The giant panda typically lives around 20 years in the wild and up to 30 years in
captivity Captivity, or being held captive, is a state wherein humans or other animals are confined to a particular space and prevented from leaving or moving freely. An example in humans is imprisonment. Prisoners of war are usually held in captivity by a ...
. A female named Jia Jia was the oldest giant panda ever in captivity, born in 1978 and died at an age of 38 on 16 October 2016.


Pathology

A seven-year-old female named Jin Yi died in 2014 in a zoo in
Zhengzhou Zhengzhou (; ), also spelt Zheng Zhou and alternatively romanized as Chengchow, is the capital and largest city of Henan Province in the central part of the People's Republic of China. Located in north-central Henan, it is one of the National ...
, China, after showing symptoms of
gastroenteritis Gastroenteritis, also known as infectious diarrhea and gastro, is an inflammation of the gastrointestinal tract including the stomach and intestine. Symptoms may include diarrhea, vomiting, and abdominal pain. Fever, lack of energy, and dehydra ...
and
respiratory disease Respiratory diseases, or lung diseases, are pathology, pathological conditions affecting the organs and tissues that make gas exchange difficult in Breathing, air-breathing animals. They include conditions of the respiratory tract including the t ...
. It was found that the cause of death was
toxoplasmosis Toxoplasmosis is a parasitic disease caused by ''Toxoplasma gondii'', an apicomplexan. Infections with toxoplasmosis are associated with a variety of neuropsychiatric and behavioral conditions. Occasionally, people may have a few weeks or months ...
, a disease caused by an obligate intracellular parasitic
protozoa Protozoa (singular: protozoan or protozoon; alternative plural: protozoans) are a group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic tissues and debris. Histo ...
n known as ''
Toxoplasma gondii ''Toxoplasma gondii'' () is an obligate intracellular parasitic protozoan (specifically an apicomplexan) that causes toxoplasmosis. Found worldwide, ''T. gondii'' is capable of infecting virtually all warm-blooded animals, but felids, such as d ...
'' that infects most warm-blooded animals, including humans.


Genomics

The giant panda
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding ge ...
was sequenced in 2009 using
Illumina dye sequencing Illumina dye sequencing is a technique used to determine the series of base pairs in DNA, also known as DNA sequencing. The reversible terminated chemistry concept was invented by Bruno Canard and Simon Sarfati at the Pasteur Institute in Paris. I ...
. Its genome contains 20 pairs of
autosome An autosome is any chromosome that is not a sex chromosome. The members of an autosome pair in a diploid cell have the same morphology, unlike those in allosome, allosomal (sex chromosome) pairs, which may have different structures. The DNA in au ...
s and one pair of sex chromosomes.


Ecology


Diet

Despite its
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
classification as a
carnivora Carnivora is a Clade, monophyletic order of Placentalia, placental mammals consisting of the most recent common ancestor of all felidae, cat-like and canidae, dog-like animals, and all descendants of that ancestor. Members of this group are f ...
n, the giant panda's
diet Diet may refer to: Food * Diet (nutrition), the sum of the food consumed by an organism or group * Dieting, the deliberate selection of food to control body weight or nutrient intake ** Diet food, foods that aid in creating a diet for weight loss ...
is primarily
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, consisting almost exclusively of bamboo. However, the giant panda still has the digestive system of a carnivore, as well as carnivore-specific genes, and thus derives little energy and little protein from consumption of bamboo. The ability to break down cellulose and lignin is very weak, and their main source of nutrients comes from starch and hemicelluloses. The most important part of their bamboo diet is the shoots, that are rich in starch which they have a higher capability of digest than strict carnivores, and have up till 32% protein content. During the shoot season, which lasts from April to August, they put on a lot of weight, which allows them to get through the nutrient-scarce period from late August to April, when they feed mostly on bamboo leaves. Pandas are born with sterile intestines and require bacteria obtained from their mother's feces to digest vegetation. The giant panda is a highly specialised animal with unique adaptations, and has lived in bamboo forests for millions of years. The average giant panda eats as much as 9 to 14 kg (20 to 30 lb) of bamboo shoots a day to compensate for the limited energy content of its diet. Ingestion of such a large quantity of material is possible and necessary because of the rapid passage of large amounts of indigestible plant material through the short, straight digestive tract. It is also noted, however, that such rapid passage of digesta limits the potential of microbial digestion in the gastrointestinal tract, limiting alternative forms of digestion. Given this voluminous diet, the giant panda defecates up to 40 times a day. The limited energy input imposed on it by its diet has affected the panda's behavior. The giant panda tends to limit its social interactions and avoids steeply sloping terrain to limit its energy expenditures. It has been estimated that an adult panda absorbs 54.8–66.1 mg of
cyanide Cyanide is a naturally occurring, rapidly acting, toxic chemical that can exist in many different forms. In chemistry, a cyanide () is a chemical compound that contains a functional group. This group, known as the cyano group, consists of ...
a day through its diet. To prevent poisoning, they have evolved anti-toxic mechanisms to protect themselves. About 80% of the cyanide is metabolized to less toxic thiocyanate and discharged in urine, while the remaining 20% is detoxified by other minor pathways. Two of the panda's most distinctive features, its large size and round face, are
adaptation In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
s to its bamboo diet. Anthropologist
Russell Ciochon Russell Ciochon (born March 11, 1948) is an American paleoanthropologist. He was born in Altadena, California and received three degrees (B.A. in 1971; M.A. in 1974; and PhD. in 1986) in anthropology from the University of California, Berkeley. He ...
observed: "
uch Uch ( pa, ; ur, ), frequently referred to as Uch Sharīf ( pa, ; ur, ; ''"Noble Uch"''), is a historic city in the southern part of Pakistan's Punjab, Pakistan, Punjab province. Uch may have been founded as Alexandria on the Indus, a town ...
like the vegetarian gorilla, the low body surface area to body volume
f the giant panda F, or f, is the sixth letter in the Latin alphabet, used in the modern English alphabet, the alphabets of other western European languages and others worldwide. Its name in English is ''ef'' (pronounced ), and the plural is ''efs''. Hist ...
is indicative of a lower metabolic rate. This lower metabolic rate and a more sedentary lifestyle allows the giant panda to subsist on nutrient poor resources such as bamboo." Similarly, the giant panda's round face is the result of powerful jaw muscles, which attach from the top of the head to the jaw. Large molars crush and grind fibrous plant material. The morphological characteristics of extinct relatives of the giant panda suggest that while the ancient giant panda was omnivorous 7 million years ago (mya), it only became herbivorous some 2–2.4 mya with the emergence of '' A. microta''. Genome sequencing of the giant panda suggests that the dietary switch could have initiated from the loss of the sole T1R1/T1R3
umami Umami ( from ja, 旨味 ), or savoriness, is one of the five basic tastes. It has been described as savory and is characteristic of broths and cooked meats. People taste umami through taste receptors that typically respond to glutamates and ...
taste receptor, resulting from two frameshift mutations within the T1R1 exons. Umami taste corresponds to high levels of glutamate as found in meat and may have thus altered the food choice of the giant panda. Although the pseudogenisation of the umami taste receptor in ''Ailuropoda'' coincides with the dietary switch to herbivory, it is likely a result of, and not the reason for, the dietary change. The mutation time for the T1R1 gene in the giant panda is estimated to 4.2 mya while fossil evidence indicates bamboo consumption in the giant panda species at least 7 mya, signifying that although complete herbivory occurred around 2 mya, the dietary switch was initiated prior to T1R1 loss-of-function. Pandas eat any of 25 bamboo species in the wild, such as ''
Fargesia dracocephala ''Fargesia dracocephala'' is a woody bamboo native to central China. It is known in Chinese as ''longtou jianzhu'' (), meaning ''dragon head Fargesia''. The scientific name is derived from Ancient Greek and has the same meaning. It is found at ...
'' and ''
Fargesia rufa ''Fargesia rufa'' is a woody bamboo native to western China. It is known in Chinese as ''qingchuan jianzhu'' (), meaning "''Qingchuan Fargesia''", ''Qingchuan'' being a county within the prefecture-level city of Guangyuan in the north of Sichuan ...
''. Only a few bamboo species are widespread at the high altitudes pandas now inhabit. Bamboo leaves contain the highest protein levels; stems have less. Because of the synchronous flowering, death, and regeneration of all bamboo within a species, the giant panda must have at least two different species available in its range to avoid starvation. While primarily herbivorous, the giant panda still retains decidedly ursine teeth and will eat meat, fish, and eggs when available. In captivity, zoos typically maintain the giant panda's bamboo diet, though some will provide specially formulated biscuits or other dietary supplements. Pandas will travel between different habitats if they need to, so they can get the nutrients that they need and to balance their diet for reproduction. For six years, scientists studied six pandas tagged with GPS collars at the Foping Reserve in the Qinling Mountains. They took note of their foraging and mating habits and analyzed samples of their food and feces. The pandas would move from the valleys into the Qinling Mountains and would only return to the valleys in autumn. During the summer months,
bamboo shoot Bamboo shoots or bamboo sprouts are the edible shoots (new bamboo culms that come out of the ground) of many bamboo species including ''Bambusa vulgaris'' and ''Phyllostachys edulis''. They are used as vegetables in numerous Asian dishes and b ...
s rich in protein are only available at higher altitudes which causes low calcium rates in the pandas. During breeding season, the pandas would return to lower altitudes to eat bamboo leaves rich in calcium.


Predators

Although adult giant pandas have few natural predators other than humans, young cubs are vulnerable to attacks by
snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a Felidae, felid in the genus ''Panthera'' native to the mountain ranges of Central Asia, Central and South Asia. It is listed as Vulnerable species, Vulnerable on the IUCN Red ...
s,
yellow-throated marten The yellow-throated marten (''Martes flavigula'') is a marten species native to Asia. It is listed as Least Concern on the IUCN Red List due to its wide distribution, evidently relatively stable population, occurrence in a number of protected are ...
s, eagles, feral dogs, and the
Asian black bear The Asian black bear (''Ursus thibetanus''), also known as the Asiatic black bear, moon bear and white-chested bear, is a medium-sized bear species native to Asia that is largely adapted to an arboreal lifestyle. It lives in the Himalayas, sout ...
. Sub-adults weighing up to may be vulnerable to predation by
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
s.


Behavior

The giant panda is a
terrestrial animal Terrestrial animals are animals that live predominantly or entirely on land (e.g. cats, dogs, ants, spiders), as compared with aquatic animals, which live predominantly or entirely in the water (e.g. fish, lobsters, octopuses), and amphibians, ...
and primarily spends its life roaming and feeding in the bamboo forests of the
Qinling Mountains The Qinling () or Qin Mountains, formerly known as the Nanshan ("Southern Mountains"), are a major east–west mountain range in southern Shaanxi Province, China. The mountains mark the divide between the drainage basins of the Yangtze and Yellow ...
and in the hilly province of
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
. Giant pandas are generally solitary. Each adult has a defined territory and a female is not tolerant of other females in her range. Social encounters occur primarily during the brief breeding season in which pandas in proximity to one another will gather. After mating, the male leaves the female alone to raise the cub. Pandas were thought to fall into the
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
category, those who are active twice a day, at dawn and dusk; however, Jindong Zhang found that pandas may belong to a category all of their own, with activity peaks in the morning, afternoon and midnight. The low nutrition quality of bamboo means pandas need to eat more frequently, and due to their lack of major predators they can be active at any time of the day. Activity is highest in June and decreases in late summer to autumn with an increase from November through the following March. Activity is also directly related to the amount of sunlight during colder days. Pandas communicate through vocalisation and scent marking such as clawing trees or spraying urine. They are able to climb and take shelter in hollow trees or rock crevices, but do not establish permanent dens. For this reason, pandas do not
hibernate Hibernation is a state of minimal activity and metabolic depression undergone by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It most ...
, which is similar to other subtropical mammals, and will instead move to elevations with warmer temperatures. Pandas rely primarily on
spatial memory In cognitive psychology and neuroscience, spatial memory is a form of memory responsible for the recording and recovery of information needed to plan a course to a location and to recall the location of an object or the occurrence of an event. Sp ...
rather than
visual memory Visual memory describes the relationship between perceptual processing and the encoding, storage and retrieval of the resulting neural representations. Visual memory occurs over a broad time range spanning from eye movements to years in order ...
. Though the panda is often assumed to be docile, it has been known to attack humans, presumably out of irritation rather than aggression. Pandas have been known to cover themselves in horse manure to protect themselves against cold temperatures.


Olfactory communication

Giant pandas heavily rely on olfactory communication to communicate with conspecifics. Scent marks are used to spread these chemical cues and are placed on landmarks like rocks or trees. Chemical communication in giant pandas plays many roles in their social situations. Scent marks and odors are used to spread information about sexual status, whether a female is in
estrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous p ...
or not, age, gender, individuality, dominance over territory, and choice of settlement. Giant pandas communicate by excreting volatile compounds, or scent marks, through the anogenital gland. These volatile compounds are found in urine and vaginal and anal secretions from the anogenital gland. The anogenital gland secretes
short chain fatty acids Short-chain fatty acids (SCFAs) are fatty acids with fewer than six carbon atoms. Derived from intestinal microbial fermentation of indigestible foods, SCFAs are the main energy source of colonocytes, making them crucial to gastrointestinal health. ...
(SCFA) and
aromatics Aromatic compounds, also known as "mono- and polycyclic aromatic hydrocarbons", are organic compounds containing one or more aromatic rings. The parent member of aromatic compounds is benzene. The word "aromatic" originates from the past grouping ...
, which are present in the scent marks of giant pandas. Giant pandas have unique positions in which they will scent mark. Males deposit scent marks or urine by lifting their hind leg, rubbing their backside, or standing in order to rub the anogenital gland onto a landmark. Females; however, exercise squatting or simply rubbing their genitals onto a landmark. The season plays a major role in mediating chemical communication. Depending on the season, mainly whether it's breeding season or not, may influence which odors are prioritized. Chemical signals can have different functions in different seasons. During the non-breeding season, females prefer the odors of other females because reproduction is not their primary motivation. However, during breeding season, odors from the opposite sex will be more attractive. Because they are solitary mammals and their breeding season is so brief, female pandas secrete chemical cues in order to let males know their sexual status. The chemical cues female pandas secrete can be considered to be
pheromones A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
for sexual reproduction. Females deposit scent marks through their urine which induces an increase in
androgen An androgen (from Greek ''andr-'', the stem of the word meaning "man") is any natural or synthetic steroid hormone that regulates the development and maintenance of male characteristics in vertebrates by binding to androgen receptors. This inc ...
levels in males. Androgen is a sex hormone found in both males and females;
testosterone Testosterone is the primary sex hormone and anabolic steroid in males. In humans, testosterone plays a key role in the development of Male reproductive system, male reproductive tissues such as testes and prostate, as well as promoting secondar ...
is the major androgen produced by males.
Civetone Civetone is a macrocyclic ketone and the main odorous constituent of civet oil. It is a pheromone sourced from the African civet. It has a strong musky odor that becomes pleasant at extreme dilutions. Civetone is closely related to muscone, the p ...
and
decanoic acid Capric acid, also known as decanoic acid or decylic acid, is a saturated fatty acid, medium-chain fatty acid (MCFA), and carboxylic acid. Its formula is CH3(CH2)8COOH. Salts and esters of decanoic acid are called caprates or decanoates. The term ...
are chemicals found in female urine which promote behavioral responses in males; both chemicals are considered giant panda pheromones. Male pandas also secrete chemical signals that include information about their sexual reproductivity and age, which is beneficial for a female when choosing a mate. For example, age can be useful for a female to determine sexual maturity and sperm quality. Pandas are also able to determine when the signal was placed, further aiding in the quest to find a potential mate. However, chemical cues are not just used for communication between males and females, pandas can determine individuality from chemical signals. This allows them to be able to differentiate between a potential partner or someone of the same sex, which could be a potential competitor. Chemical cues, or odors, play an important role in how a panda choses their habitat. Pandas look for conspecific odors that tell them not only the identity of another panda, but if they should avoid them or not. Pandas tend to avoid their species for most of the year, breeding season being the brief time of major interaction. Chemical signaling allows for avoidance and
competition Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indivi ...
. Pandas whose habitats are in similar locations will collectively leave scent marks in a unique location which is termed "scent stations." When pandas come across these scent stations, they are able to identify a specific panda and the scope of their habitat. This allows pandas to be able to pursue a potential mate or avoid a potential competitor. Pandas can assess an individuals dominance status, including their age and size, via odor cues and may chose to avoid a scent mark if the signaler's competitive ability outweighs their own. A pandas size can be conveyed through the height of the scent mark. Since larger animals can place higher scent marks, an elevated scent mark advertises a higher competitive ability. Age must also be taken into consideration when assessing a competitor's fighting ability. For example, a mature panda will be larger than a younger, immature panda and posses an advantage during a fight.


Reproduction

Initially, the primary method of breeding giant pandas in captivity was by
artificial insemination Artificial insemination is the deliberate introduction of sperm into a female's cervix or uterine cavity for the purpose of achieving a pregnancy through in vivo fertilization by means other than sexual intercourse. It is a fertility treatme ...
, as they seemed to lose their interest in
mating In biology, mating is the pairing of either opposite-sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reproduc ...
once they were captured. This led some scientists to try extreme methods, such as showing them videos of giant pandas mating and giving the males
sildenafil Sildenafil, sold under the brand name Viagra, among others, is a medication used to treat erectile dysfunction and pulmonary arterial hypertension. It is unclear if it is effective for treating sexual dysfunction in women. It is taken by ...
(commonly known as Viagra). Only recently have researchers started having success with captive breeding programs, and they have now determined giant pandas have comparable breeding to some populations of the
American black bear The American black bear (''Ursus americanus''), also called simply a black bear or sometimes a baribal, is a medium-sized bear endemic to North America. It is the continent's smallest and most widely distributed bear species. American black bear ...
, a thriving bear species. The normal reproductive rate is considered to be one young every two years. Giant pandas reach sexual maturity between the ages of four and eight, and may be reproductive until age 20. The mating season is between March and May, when a female goes into
estrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous p ...
, which lasts for two or three days and only occurs once a year. When mating, the female is in a crouching, head-down position as the male mounts her from behind.
Copulation Sexual intercourse (or coitus or copulation) is a sexual activity typically involving the insertion and thrusting of the penis into the vagina for sexual pleasure or reproduction.Sexual intercourse most commonly means penile–vaginal penetrat ...
time ranges from 30 seconds to five minutes, but the male may mount her repeatedly to ensure successful fertilisation. The
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
period is somewhere between 95 and 160 days - the variability is due to the fact that the fertilized egg may linger in the reproductive system for a while before implanting on the uterine wall. Giant pandas give birth to twins in about half of pregnancies. If twins are born, usually only one survives in the wild. The mother will select the stronger of the cubs, and the weaker cub will die due to starvation. The mother is thought to be unable to produce enough milk for two cubs since she does not store fat. The father has no part in helping raise the cub. When the cub is first born, it is pink, blind, and toothless, weighing only , or about of the mother's weight, proportionally the smallest baby of any placental mammal. It nurses from its mother's breast six to 14 times a day for up to 30 minutes at a time. For three to four hours, the mother may leave the den to feed, which leaves the cub defenseless. One to two weeks after birth, the cub's skin turns grey where its hair will eventually become black. Slight pink colour may appear on the cub's fur, as a result of a
chemical reaction A chemical reaction is a process that leads to the IUPAC nomenclature for organic transformations, chemical transformation of one set of chemical substances to another. Classically, chemical reactions encompass changes that only involve the pos ...
between the fur and its mother's
saliva Saliva (commonly referred to as spit) is an extracellular fluid produced and secreted by salivary glands in the mouth. In humans, saliva is around 99% water, plus electrolytes, mucus, white blood cells, epithelial cells (from which DNA can be ...
. A month after birth, the colour pattern of the cub's fur is fully developed. Its fur is very soft and coarsens with age. The cub begins to crawl at 75 to 80 days; mothers play with their cubs by rolling and wrestling with them. The cubs can eat small quantities of bamboo after six months, though mother's milk remains the primary food source for most of the first year. Giant panda cubs weigh 45 kg (100 pounds) at one year and live with their mothers until they are 18 months to two years old. The interval between births in the wild is generally two years. In July 2009, Chinese scientists confirmed the birth of the first cub to be successfully conceived through artificial insemination using frozen sperm. The cub was born at 07:41 on 23 July that year in
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
as the third cub of You You, an 11-year-old. The technique for freezing the sperm in
liquid nitrogen Liquid nitrogen—LN2—is nitrogen in a liquid state at low temperature. Liquid nitrogen has a boiling point of about . It is produced industrially by fractional distillation of liquid air. It is a colorless, low viscosity liquid that is wide ...
was first developed in 1980 and the first birth was hailed as a solution to the dwindling availability of giant panda semen, which had led to inbreeding. Panda semen, which can be frozen for decades, could be shared between different zoos to save the species. It is expected that zoos in destinations such as San Diego in the United States and
Mexico City Mexico City ( es, link=no, Ciudad de México, ; abbr.: CDMX; Nahuatl: ''Altepetl Mexico'') is the capital and largest city of Mexico, and the most populous city in North America. One of the world's alpha cities, it is located in the Valley o ...
will now be able to provide their own semen to inseminate more giant pandas. In August 2014, a rare birth of panda triplets was announced in China; it was the fourth of such births ever reported. Attempts have also been made to reproduce giant pandas by
interspecific pregnancy Interspecific pregnancy (literally ''pregnancy between species'', also called interspecies pregnancy or xenopregnancy)Page 126 in: is the pregnancy involving an embryo or fetus belonging to another species than the carrier. Strictly, it excludes ...
where cloned panda embryos were implanted into the uterus of an animal of another species. This has resulted in panda fetuses, but no live births.


Human use and interaction


Early references

In the past, pandas were thought to be rare and noble creatures – the
Empress Dowager Bo Empress Dowager Bo (薄太后) was an imperial concubine of Emperor Gaozu of Han (Liu Bang). She was also known as Consort Bo (薄姬) during the life of the Emperor, and more formally as either Empress Dowager Xiaowen (孝文太后) or (rarer) E ...
was buried with a panda skull in her vault. The grandson of
Emperor Taizong of Tang Emperor Taizong of Tang (28January 59810July 649), previously Prince of Qin, personal name Li Shimin, was the second emperor of the Tang dynasty of China, ruling from 626 to 649. He is traditionally regarded as a co-founder of the dynasty ...
is said to have given Japan two pandas and a sheet of panda skin as a sign of goodwill. Unlike many other animals in
Ancient China The earliest known written records of the history of China date from as early as 1250 BC, from the Shang dynasty (c. 1600–1046 BC), during the reign of king Wu Ding. Ancient historical texts such as the '' Book of Documents'' (early chapte ...
, pandas were rarely thought to have medical uses. The few known uses include the Sichuan tribal peoples' use of panda urine to melt accidentally swallowed needles, and the use of panda pelts to control
menstruation Menstruation (also known as a period, among other colloquial terms) is the regular discharge of blood and mucosal tissue from the inner lining of the uterus through the vagina. The menstrual cycle is characterized by the rise and fall of hor ...
as described in the
Qin dynasty The Qin dynasty ( ; zh, c=秦朝, p=Qín cháo, w=), or Ch'in dynasty in Wade–Giles romanization ( zh, c=, p=, w=Ch'in ch'ao), was the first Dynasties in Chinese history, dynasty of Imperial China. Named for its heartland in Qin (state), ...
encyclopedia ''
Erya The ''Erya'' or ''Erh-ya'' is the first surviving Chinese dictionary. Bernhard Karlgren (1931:49) concluded that "the major part of its glosses must reasonably date from the 3rd century BC." Title Chinese scholars interpret the first title chara ...
''. The creature named ''mo'' (貘) mentioned in some ancient books has been interpreted as giant panda. The dictionary ''
Shuowen Jiezi ''Shuowen Jiezi'' () is an ancient Chinese dictionary from the Han dynasty. Although not the first comprehensive Chinese character dictionary (the '' Erya'' predates it), it was the first to analyze the structure of the characters and to give ...
'' (
Eastern Han Dynasty The Han dynasty (, ; ) was an imperial dynasty of China (202 BC – 9 AD, 25–220 AD), established by Liu Bang (Emperor Gao) and ruled by the House of Liu. The dynasty was preceded by the short-lived Qin dynasty (221–207 BC) and a warr ...
) says that the ''mo'', from Shu (Sichuan), is bear-like, but yellow-and-black, although the older ''
Erya The ''Erya'' or ''Erh-ya'' is the first surviving Chinese dictionary. Bernhard Karlgren (1931:49) concluded that "the major part of its glosses must reasonably date from the 3rd century BC." Title Chinese scholars interpret the first title chara ...
'' describes ''mo'' simply as a "white leopard". The interpretation of the legendary fierce creature ''
pixiu Pixiu (; Old Chinese ( ZS) *''bi-qʰu'') is a Chinese mythical hybrid creature. Pixiu are considered powerful protectors of feng shui practitioners, and resemble strong, winged lions. A Pixiu is an earth and sea variation, particularly an infl ...
'' (貔貅) as referring to the giant panda is also common. During the reign of the
Yongle Emperor The Yongle Emperor (; pronounced ; 2 May 1360 – 12 August 1424), personal name Zhu Di (), was the third Emperor of the Ming dynasty, reigning from 1402 to 1424. Zhu Di was the fourth son of the Hongwu Emperor, the founder of the Ming dyn ...
(early 15th century), his relative from
Kaifeng Kaifeng () is a prefecture-level city in east-central Henan province, China. It is one of the Eight Ancient Capitals of China, having been the capital eight times in history, and is best known for having been the Chinese capital during the Nort ...
sent him a captured ''
zouyu Zouyu (), also called ''zouwu'' () or ''zouya'' (), is a legendary creature mentioned in old Chinese literature. The earliest known appearance of the characters (''zou yu'') is in the '' Book of Songs'', but J.J.L. Duyvendak describes that the ...
'' ( 騶虞), and another ''zouyu'' was sighted in
Shandong Shandong ( , ; ; alternately romanized as Shantung) is a coastal province of the People's Republic of China and is part of the East China region. Shandong has played a major role in Chinese history since the beginning of Chinese civilizati ...
. ''Zouyu'' is a legendary "righteous" animal, which, similarly to a ''
qilin The qilin (; ) is a legendary hooved chimerical creature that appears in Chinese mythology, and is said to appear with the imminent arrival or passing of a sage or illustrious ruler. Qilin are a specific type of the mythological family of on ...
'', only appears during the rule of a benevolent and sincere monarch. It is said to be fierce as a tiger, but gentle and strictly vegetarian, and described in some books as a
white tiger The white tiger or bleached tiger is a leucistic pigmentation variant of the Mainland tiger. It is reported in the wild from time to time in the Indian states of Madhya Pradesh, Assam, West Bengal, Bihar, Odisha, in the Sunderbans region and ...
with black spots. Puzzled about the real zoological identity of the creature captured during the Yongle era, Dutch Sinologist
J. J. L. Duyvendak Jan Julius Lodewijk Duyvendak (28 June 18899 July 1954) was a Dutch Sinologist and professor of Chinese at Leiden University. He is known for his translation of ''The Book of Lord Shang'' and his studies of the ''Dao De Jing''. He was co-editor ...
exclaimed, "Can it possibly have been a Pandah?"


Western discovery

The West first learned of the giant panda on 11 March 1869, when the French
missionary A missionary is a member of a Religious denomination, religious group which is sent into an area in order to promote its faith or provide services to people, such as education, literacy, social justice, health care, and economic development.Tho ...
Armand David Father Armand David (7 September 1826, Espelette – 10 November 1900, Paris) was a Lazarist missionary Catholic priest as well as a zoologist and a botanist. Several species, such as Père David's deer, are named after him — be ...
received a skin from a hunter. The first Westerner known to have seen a living giant panda is the German zoologist
Hugo Weigold Max Hugo Weigold (27 May 1886 – 9 July 1973) was a German zoologist and a pioneer bird bander who worked at the Heligoland Bird Observatory, one of the world's first bird-ringing sites. Weigold was born in Dresden. He studied natural sciences ...
, who purchased a cub in 1916. Kermit and
Theodore Roosevelt, Jr. Theodore Roosevelt III ( ), often known as Theodore Jr.Morris, Edmund (1979). ''The Rise of Theodore Roosevelt''. index.While it was President Theodore Roosevelt who was legally named Theodore Roosevelt Jr., the President's fame made it simple ...
, became the first Westerners to shoot a panda, on an expedition funded by the
Field Museum of Natural History The Field Museum of Natural History (FMNH), also known as The Field Museum, is a natural history museum in Chicago, Illinois, and is one of the largest such museums in the world. The museum is popular for the size and quality of its educational ...
in the 1920s. In 1936,
Ruth Harkness Ruth Elizabeth Harkness (21 September 1900 - 20 July 1947) was an American fashion designer and socialite who traveled to China in 1936 and brought out the first live giant panda to the United States - not in a cage or on a leash, but wrapped in h ...
became the first Westerner to bring back a live giant panda, a cub named Su Lin who went to live at the
Brookfield Zoo Brookfield Zoo, also known as the Chicago Zoological Park, is a zoo located in the Chicago suburb of Brookfield, Illinois. It houses around 450 species of animals in an area of . It opened on July 1, 1934, and quickly gained international recogni ...
in Chicago. In 1938, Floyd Tangier Smith captured and delivered five giant pandas to London, they arrived on 23 December aboard the SS ''Antenor''. These five were the first on British soil and were transferred to
London Zoo London Zoo, also known as ZSL London Zoo or London Zoological Gardens is the world's oldest scientific zoo. It was opened in London on 27 April 1828, and was originally intended to be used as a collection for science, scientific study. In 1831 o ...
. One, named Grandma, only lasted a few days. She was taxidermized by E. Gerrard and Sons and sold to
Leeds City Museum Leeds City Museum, originally established in 1819, reopened in 2008 in Leeds, West Yorkshire, England. It is housed in the former Mechanics' Institute built by Cuthbert Brodrick, in Cookridge Street (now Millennium Square). It is one of nine s ...
where she is currently on display to the public. Another, Ming, became
London Zoo London Zoo, also known as ZSL London Zoo or London Zoological Gardens is the world's oldest scientific zoo. It was opened in London on 27 April 1828, and was originally intended to be used as a collection for science, scientific study. In 1831 o ...
's first Giant Panda. Her skull is held by the
Royal College of Surgeons The Royal College of Surgeons is an ancient college (a form of corporation) established in England to regulate the activity of surgeons. Derivative organisations survive in many present and former members of the Commonwealth. These organisations a ...
of England.


Panda diplomacy

In the 1970s, gifts of giant pandas to American and Japanese zoos formed an important part of the diplomacy of the People's Republic of China (PRC), as it marked some of the first cultural exchanges between China and the West. This practice has been termed "panda diplomacy". By 1984, however, pandas were no longer given as gifts. Instead, China began to offer pandas to other nations only on 10-year loans, under terms including a fee of up to US$1,000,000 per year and a provision that any cubs born during the loan are the property of China. Since 1998, because of a WWF
lawsuit - A lawsuit is a proceeding by a party or parties against another in the civil court of law. The archaic term "suit in law" is found in only a small number of laws still in effect today. The term "lawsuit" is used in reference to a civil actio ...
, the
United States Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
only allows a US
zoo A zoo (short for zoological garden; also called an animal park or menagerie) is a facility in which animals are kept within enclosures for public exhibition and often bred for Conservation biology, conservation purposes. The term ''zoological g ...
to import a panda if the zoo can ensure that China will channel more than half of its loan fee into
conservation Conservation is the preservation or efficient use of resources, or the conservation of various quantities under physical laws. Conservation may also refer to: Environment and natural resources * Nature conservation, the protection and manageme ...
efforts for the giant panda and its habitat. As a result of this change in policy, nearly all the pandas in the world are owned by China. The pandas leased to foreign zoos and any cubs are eventually returned to China. In May 2005, China offered a breeding pair to
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
. The issue became embroiled in
cross-Strait relations Cross-Strait relations (sometimes called Mainland–Taiwan relations, or Taiwan-China relations) are the relations between China (officially the People's Republic of China) and Taiwan (officially the Republic of China). The relationship ...
– both over the underlying symbolism, and over technical issues such as whether the transfer would be considered "domestic" or "international", or whether any true conservation purpose would be served by the exchange. A contest in 2006 to name the pandas was held in the mainland, resulting in the politically charged names
Tuan Tuan and Yuan Yuan Tuan Tuan and Yuan Yuan were two giant pandas that were gifted by the People's Republic of China (PRC, mainland China) to the Republic of China (ROC, Taiwan) in 2008 as part of a cultural exchange program. The idea was first proposed in 2005, but ...
(from ''tuanyuan'', meaning "reunion", i.e. "reunification"). China's offer was initially rejected by
Chen Shui-bian Chen Shui-bian (; born 12 October 1950) is a retired Taiwanese politician and lawyer who served as the president of the Republic of China (Taiwan) from 2000 to 2008. Chen was the first president from the Democratic Progressive Party (DPP) whic ...
, then President of Taiwan. However, when
Ma Ying-jeou Ma Ying-jeou ( zh, 馬英九, born 13 July 1950) is a Hong Kong-born Taiwanese politician who served as president of the Republic of China from 2008 to 2016. Previously, he served as justice minister from 1993 to 1996 and mayor of Taipei from ...
assumed the presidency in 2008, the offer was accepted, and the pandas arrived in December of that year.


Zoos

Pandas have been kept in zoos as early as the
Western Han Dynasty The Han dynasty (, ; ) was an Dynasties in Chinese history, imperial dynasty of China (202 BC – 9 AD, 25–220 AD), established by Emperor Gaozu of Han, Liu Bang (Emperor Gao) and ruled by the House of Liu. The dynasty was preceded by th ...
in China, where the writer
Sima Xiangru Sima Xiangru ( , ; c. 179117BC) was a Chinese musician, poet, and politician who lived during the Western Han dynasty. Sima is a significant figure in the history of Classical Chinese poetry, and is generally regarded as the greatest of all com ...
noted that the panda was the most treasured animal in the emperor's garden of exotic animals in the capital
Chang'an Chang'an (; ) is the traditional name of Xi'an. The site had been settled since Neolithic times, during which the Yangshao culture was established in Banpo, in the city's suburbs. Furthermore, in the northern vicinity of modern Xi'an, Qin Shi ...
(present
Xi'an Xi'an ( , ; ; Chinese: ), frequently spelled as Xian and also known by #Name, other names, is the list of capitals in China, capital of Shaanxi, Shaanxi Province. A Sub-provincial division#Sub-provincial municipalities, sub-provincial city o ...
). Not until the 1950s were pandas again recorded to have been exhibited in China's zoos. Chi Chi at the
London Zoo London Zoo, also known as ZSL London Zoo or London Zoological Gardens is the world's oldest scientific zoo. It was opened in London on 27 April 1828, and was originally intended to be used as a collection for science, scientific study. In 1831 o ...
became very popular. This influenced the
World Wildlife Fund The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wo ...
to use a panda as its symbol. A 2006 ''
New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid d ...
'' article outlined the economics of keeping pandas, which costs five times more than keeping the next most expensive animal, an elephant. American zoos generally pay the Chinese government $1 million a year in fees, as part of a typical ten-year contract. San Diego's contract with China was to expire in 2008, but got a five-year extension at about half of the previous yearly cost. The last contract, with the
Memphis Zoo The Memphis Zoo, located in Midtown, Memphis, Tennessee, United States, is home to more than 3,500 animals representing over 500 different species. Created in April 1906, the zoo has been a major tenant of Overton Park for more than 100 years. Th ...
in
Memphis, Tennessee Memphis is a city in the U.S. state of Tennessee. It is the seat of Shelby County in the southwest part of the state; it is situated along the Mississippi River. With a population of 633,104 at the 2020 U.S. census, Memphis is the second-mos ...
, ended in 2013.


Conservation

The giant panda is a
vulnerable species A vulnerable species is a species which has been Conservation status, categorized by the International Union for Conservation of Nature as being threatened species, threatened with extinction unless the circumstances that are threatened species, ...
, threatened by continued
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
and
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological processes ...
, and by a very low birthrate, both in the wild and in
captivity Captivity, or being held captive, is a state wherein humans or other animals are confined to a particular space and prevented from leaving or moving freely. An example in humans is imprisonment. Prisoners of war are usually held in captivity by a ...
. Its range is currently confined to a small portion on the western edge of its historical range, which stretched through southern and eastern China, northern
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, and northern
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
. The giant panda has been a target of poaching by locals since ancient times and by foreigners since it was introduced to the West. Starting in the 1930s, foreigners were unable to poach giant pandas in China because of the
Second Sino-Japanese War The Second Sino-Japanese War (1937–1945) or War of Resistance (Chinese term) was a military conflict that was primarily waged between the Republic of China and the Empire of Japan. The war made up the Chinese theater of the wider Pacific Th ...
and the
Chinese Civil War The Chinese Civil War was fought between the Kuomintang-led government of the Republic of China and forces of the Chinese Communist Party, continuing intermittently since 1 August 1927 until 7 December 1949 with a Communist victory on m ...
, but pandas remained a source of soft furs for the locals. The population boom in China after 1949 created stress on the pandas' habitat and the subsequent famines led to the increased hunting of wildlife, including pandas. During the
Cultural Revolution The Cultural Revolution, formally known as the Great Proletarian Cultural Revolution, was a sociopolitical movement in the People's Republic of China (PRC) launched by Mao Zedong in 1966, and lasting until his death in 1976. Its stated goal ...
, all studies and conservation activities on the pandas were stopped. After the
Chinese economic reform The Chinese economic reform or reform and opening-up (), known in the West as the opening of China, is the program of economic reforms termed " Socialism with Chinese characteristics" and "socialist market economy" in the People's Republic of C ...
, demand for panda skins from Hong Kong and Japan led to illegal poaching for the
black market A black market, underground economy, or shadow economy is a clandestine market or series of transactions that has some aspect of illegality or is characterized by noncompliance with an institutional set of rules. If the rule defines the se ...
, acts generally ignored by the local officials at the time. In 1963, the PRC government set up
Wolong National Nature Reserve Wolong National Nature Reserve (), also known as Wolong Special Administrative Region (), is a protected area located in Wenchuan County, Sichuan Province, People's Republic of China. Established in 1963 with an initial size of about 20,000 hecta ...
to save the declining panda population. In 2006, scientists reported that the number of pandas living in the wild may have been underestimated at about 1,000. Previous population surveys had used conventional methods to estimate the size of the wild panda population, but using a new method that analyzes DNA from panda
droppings Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a relati ...
, scientists believe the wild population may be as large as 3,000. In 2006, there were 40 panda reserves in China, compared to just 13 reserves in 1998. As the species has been reclassified to "vulnerable" since 2016, the conservation efforts are thought to be working. Furthermore, in response to this reclassification, the
State Forestry Administration of China The State Forestry and Grassland Administration () is an administration of the People's Republic of China, in charge of the national forestry affairs. It was founded in 1949 as Ministry of Forestry and Farming. On November 5, 1951, it changed to ...
announced that they would not accordingly lower the conservation level for panda, and would instead reinforce the conservation efforts. The giant panda is among the world's most adored and protected rare animals, and is one of the few in the world whose natural inhabitant status was able to gain a
UNESCO World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
designation. The
Sichuan Giant Panda Sanctuaries The Sichuan Giant Panda Sanctuaries () located in southwest Sichuan province of China, is the home to more than 30% of the world's giant pandas and is among the most important sites for the captive breeding of these pandas. It covers 9245 km2 ...
, located in the southwest province of
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
and covering seven natural reserves, were inscribed onto the World Heritage List in 2006. Not all conservationists agree that the money spent on conserving pandas is well spent.
Chris Packham Christopher Gary Packham CBE (born 4 May 1961) is an English naturalist, nature photographer, television presenter and author, best known for his television work including the CBBC children's nature series ''The Really Wild Show'' from 1986 t ...
has argued that the breeding of pandas in captivity is "pointless" because "there is not enough habitat left to sustain them".Chris Packham: 'Giant pandas should be allowed to die out'
''
The Daily Telegraph ''The Daily Telegraph'', known online and elsewhere as ''The Telegraph'', is a national British daily broadsheet newspaper published in London by Telegraph Media Group and distributed across the United Kingdom and internationally. It was fo ...
''. 22 September 2009.
Packham argues that the money spent on pandas would be better spent elsewhere, and has said he would "eat the last panda if I could have all the money we have spent on panda conservation put back on the table for me to do more sensible things with". He also quoted, "The panda is possibly one of the grossest wastes of conservation money in the last half century",Beyond cute and cuddly
. ''
The Australian ''The Australian'', with its Saturday edition, ''The Weekend Australian'', is a broadsheet newspaper published by News Corp Australia since 14 July 1964.Bruns, Axel. "3.1. The active audience: Transforming journalism from gatekeeping to gatew ...
''. 10 November 2007.
though he has apologised for upsetting people who like pandas. However, a 2015 paper found that the giant panda can serve as an
umbrella species Umbrella species are species selected for making wildlife conservation, conservation-related decisions, typically because protecting these species indirectly protects the many other species that make up the ecological community (ecology), community ...
as the preservation of their habitat also helps other
endemic species Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
in China, including 70% of the country's forest birds, 70% of mammals and 31% of amphibians. In 2012,
Earthwatch Institute Earthwatch Institute is an international environmental charity. Founded as Educational Expeditions International in 1971 by Bob Citron and Clarence Truesdale, Earthwatch supports hundreds of Ph.D. researchers across dozens of countries, conduc ...
, a global nonprofit that teams volunteers with scientists to conduct important environmental research, launched a program called "On the Trail of Giant Panda". This program, based in the Wolong National Nature Reserve, allows volunteers to work up close with pandas cared for in captivity, and help them adapt to life in the wild, so that they may breed, and live longer and healthier lives. Efforts to preserve the panda bear populations in China have come at the expense of other animals in the region, including snow leopards, wolves, and dholes. In order to improve living and mating conditions for the fragmented populations of pandas, nearly 70 natural reserves have been combined to form the
Giant Panda National Park Giant Panda National Park is a large protected area within the Sichuan, Gansu, and Shaanxi provinces of China. Established in 2020, the park is made up of roughly 70 nature reserves and surpasses 10,000 square miles in size, making it more than th ...
in 2020. With a size of 10,500 square miles, the park is roughly three times as large as
Yellowstone National Park Yellowstone National Park is an American national park located in the western United States, largely in the northwest corner of Wyoming and extending into Montana and Idaho. It was established by the 42nd U.S. Congress with the Yellowston ...
and incorporates the
Wolong National Nature Reserve Wolong National Nature Reserve (), also known as Wolong Special Administrative Region (), is a protected area located in Wenchuan County, Sichuan Province, People's Republic of China. Established in 1963 with an initial size of about 20,000 hecta ...
. The state-owned
Bank of China The Bank of China (BOC; ) is a Chinese majority state-owned commercial bank headquartered in Beijing and the fourth largest bank in the world. The Bank of China was founded in 1912 by the Republican government as China's central bank, repl ...
helped to enable the project with US$1.5 billion. One major aim is to permanently keep the panda population stable enough to avoid a relapse to its former
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
"endangered" status. Especially small, isolated populations run the risk of inbreeding and smaller genetic variety makes the individuals more vulnerable to various defects and genetic
mutation In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, mi ...
. Allowing a larger group of individuals to roam through a larger area freely and choose from a greater variety of mates, helps to enrich genetic diversity of their offspring. In 2020, the panda population of the new national park was already above 1,800 individuals, which is roughly 80 percent of the entire panda population in China. Establishing the new protected area in the
Sichuan Province Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
also gives various other endangered or threatened species, like the
Siberian tiger The Siberian tiger or Amur tiger is a population of the tiger subspecies ''Panthera tigris tigris'' native to the Russian Far East, Northeast China and possibly North Korea. It once ranged throughout the Korean Peninsula, but currently inhabit ...
, the possibility to improve their living conditions by offering them a habitat. Other species who benefit from the protection of their habitat include the
snow leopard The snow leopard (''Panthera uncia''), also known as the ounce, is a Felidae, felid in the genus ''Panthera'' native to the mountain ranges of Central Asia, Central and South Asia. It is listed as Vulnerable species, Vulnerable on the IUCN Red ...
, the
golden snub-nosed monkey The golden snub-nosed monkey (''Rhinopithecus roxellana'') is an Old World monkey in the subfamily Colobinae. It is endemic to a small area in temperate, mountainous forests of central and Southwest China. They inhabit these mountainous forests ...
, the
red panda The red panda (''Ailurus fulgens''), also known as the lesser panda, is a small mammal native to the eastern Himalayas and southwestern China. It has dense reddish-brown fur with a black belly and legs, white-lined ears, a mostly white muzzle ...
and the
complex-toothed flying squirrel The complex-toothed flying squirrel (''Trogopterus xanthipes'') occurs in the southern Chinese provinces Hubei, Hunan, Guizhou, Sichuan, and Yunnan. The common name refers to the teeth, which differ from those of other species of flying squirrel ...
. In July 2021, Chinese conservation authorities announced that giant pandas are no longer endangered in the wild following years of conservation efforts, with a population in the wild exceeding 1,800.


Biofuel

Microbes in panda waste are being investigated for their use in creating
biofuels Biofuel is a fuel that is produced over a short time span from biomass, rather than by the very slow natural processes involved in the formation of fossil fuels, such as oil. According to the United States Energy Information Administration (E ...
from bamboo and other plant materials.


Population chart


See also

* Giant pandas around the world *
List of giant pandas This is a list of giant pandas, both alive and deceased. The giant panda is a conservation-reliant vulnerable species. Wild population estimates vary; one estimate shows that there are about 1,590 individuals living in the wild, while a 2006 stud ...
*
Panda tea Panda tea (Chinese: 熊猫茶), or panda dung tea, produced in the Ya’an mountainous region of Sichuan, China, is a type of tea fertilized by the dung of pandas. When it officially went on the market in April 2012, it was reputedly the world' ...
*
Pygmy giant panda ''Ailuropoda microta'' is the earliest known ancestor of the giant panda. It measured 1 m (3 ft) in length; the modern giant panda grows to a size in excess of 1.5 m (5 ft). Wear patterns on its teeth suggest it lived on a die ...
*
Wildlife of China China's vast and diverse landscape is home to a profound variety and abundance of wildlife. As of one of 17 megadiverse countries in the world, China has, according to one measure, 7,516 species of vertebrates including 4,936 fish, 1,269 bird, ...
*
List of endangered and protected species of China The endangered species of China may include any wildlife species designated for protection by the national government of China or listed as endangered by international organizations such as the Convention on International Trade in Endangered Sp ...


References

;Notes ;Bibliography * AFP (via
Discovery Channel Discovery Channel (known as The Discovery Channel from 1985 to 1995, and often referred to as simply Discovery) is an American cable channel owned by Warner Bros. Discovery, a publicly traded company run by CEO David Zaslav. , Discovery Channe ...
) (20 June 2006)
Panda Numbers Exceed Expectations
* Associated Press (via
CNN CNN (Cable News Network) is a multinational cable news channel headquartered in Atlanta, Georgia, U.S. Founded in 1980 by American media proprietor Ted Turner and Reese Schonfeld as a 24-hour cable news channel, and presently owned by the M ...
) (2006)
Article link
* Catton, Chris (1990). ''Pandas''. Christopher Helm. * Friends of the National Zoo (2006). ''Panda Cam: A Nation Watches Tai Shan the Panda Cub Grow''. New York: Fireside Books. * Goodman, Brenda (12 February 2006)

''
The New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid ...
''. * (An earlier edition is available as ''The Smithsonian Book of Giant Pandas'', Smithsonian Institution Press, 2002, .)
Panda Facts At a Glance
(N.d.). ''www.wwfchina.org''. WWF China. * Ryder, Joanne (2001). ''Little panda: The World Welcomes Hua Mei at the San Diego Zoo''. New York: Simon & Schuster. * (There are also several later reprints) * * Warren, Lynne (July 2006). "Panda, Inc." ''National Geographic''. (About Mei Xiang, Tai Shan and the Wolong Panda Research Facility in Chengdu China).


External links


BBC Nature:
Giant panda news, and video clips from BBC programmes past and present.

* ttps://web.archive.org/web/20080704204350/http://www.panda.org/about_wwf/what_we_do/species/our_solutions/endangered_species/giant_panda/index.cfm WWF– environmental conservation organization
Pandas International
– panda conservation group
National Zoo Live Panda Cams
– Baby Panda Tai Shan and mother Mei Xiang
NPR News 2007/08/20 – Panda Romance Stems From Bamboo
* View th
panda genome
on
Ensembl Ensembl genome database project is a scientific project at the European Bioinformatics Institute, which provides a centralized resource for geneticists, molecular biologists and other researchers studying the genomes of our own species and other v ...
.
Texts and pictures of the Panda exhibition at the Museum für Naturkunde Berlin

iPanda-50: annotated image dataset for fine-grained panda identification on Github
{{Authority control Mammals of China Endemic fauna of China Clawed herbivores Herbivorous mammals EDGE species Vulnerable animals Vulnerable fauna of Asia Articles containing video clips Species endangered by agricultural development Species endangered by logging Mammals described in 1869 Taxa named by Armand David