Pampatheriidae ("

Pampas The Pampas (from the qu, pampa, meaning "plain") are fertile South American low grasslands that cover more than and include the Argentine provinces of Buenos Aires, La Pampa, Santa Fe, Entre Ríos, and Córdoba; all of Uruguay; and Brazi ...

beasts") is an extinct family of large plantigrade

armored Armour (British English) or armor (American English; see spelling differences) is a covering used to protect an object, individual, or vehicle from physical injury or damage, especially direct contact weapons or projectiles during combat, or f ...

armadillos related to extant armadillos in the order

Cingulata Cingulata, part of the superorder Xenarthra, is an order of armored New World placental mammals. Dasypodids and chlamyphorids, the armadillos, are the only surviving families in the order. Two groups of cingulates much larger than extant ar ...

. However, pampatheriids have existed as a separate lineage since at least the middle

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...

Mustersan The Mustersan age is a period of geologic time (48.0–42.0 Ma) within the Eocene epoch of the Paleogene, used more specifically within the South American land mammal age (SALMA) classification. It follows the Casamayoran and precedes the Divisader ...

age, . Pampatheres evolved in

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...

during its long period of Cenozoic isolation. Although widespread, they were less diverse and abundant than the armadillos. ''

Holmesina ''Holmesina'' is a genus of pampathere, an extinct group of armadillo-like creatures that were distantly related to extant armadillos. Like armadillos, and unlike the other extinct branch of megafaunal cingulates, the glyptodonts, the shell wa ...

'' spread to North America after the formation of the Isthmus of Panama as part of the Great American Interchange. They finally disappeared on both continents in the end-Pleistocene extinctions, about 12,000 years ago.

Description

Pampatheres are believed to have attained a weight of up to . Like three-banded armadillos, and unlike glyptodonts, their armored shell was given some flexibility by three movable lateral bands of

scute A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior po ...

s. The

osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinc ...

s (bony plates in the skin comprising the armor) of pampatheres were each covered by a single

keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, ho ...

ized scute, unlike osteoderms of armadillos, which have more than one scute. Holmesina 3 Clean

A study of pampathere jaw biomechanics showed that their masticatory musculature was more powerful and more adapted for transverse movements than that of armadillos, leading to the conclusion that much of their diet was coarse vegetation. They are thought to have been primarily grazers, unlike armadillos, which are omnivorous or insectivorous. The variation between species in the expression of adaptations for grinding coarse vegetation correlates with the aridity of their habitat; such adaptations are most pronounced in '' Pampatherium typum'', which lived in the arid

, and least pronounced in ''

Holmesina occidentalis ''Holmesina'' is a genus of pampathere, an extinct group of armadillo-like creatures that were distantly related to extant armadillos. Like armadillos, and unlike the other extinct branch of megafaunal cingulates, the glyptodonts, the shell was ...

'', which lived in humid lowlands.

References

External links

Paleodatabase
† Prehistoric cingulates Oligocene first appearances Holocene extinctions Paleogene mammals of South America Neogene mammals of South America Pleistocene mammals of South America Pleistocene mammals of North America Prehistoric mammal families {{Paleo-mammal-stub