Palpus Of Scelidomachus Socotranus
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Pedipalps (commonly shortened to palps or palpi) are the second pair of
appendage An appendage (or outgrowth) is an external body part, or natural prolongation, that protrudes from an organism's body. In arthropods, an appendage refers to any of the homologous body parts that may extend from a body segment, including anten ...
s of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") and anterior to the first pair of walking legs.


Overview

Pedipalps are composed of six segments or articles: the coxa, the trochanter, the femur, the short patella, the tibia, and the tarsus. In spiders, the coxae frequently have extensions called
maxillae The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
or gnathobases, which function as mouth parts with or without some contribution from the coxae of the anterior
legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element ...
. The limbs themselves may be simple tactile organs outwardly resembling the legs, as in spiders, or chelate weapons ( pincers) of great size, as in scorpions. The pedipalps of Solifugae are covered in setae, but have not been studied in detail. Comparative studies of pedipalpal morphology may suggest that leg-like pedipalps are primitive in arachnids. At present, the only reasonable alternative to this view is to assume that Xiphosurans reflect the morphology of the primitive arachnid pedipalp and to conclude that this appendage is primitively chelate. Pedipalps are traditionally thought to be
homologous Homology may refer to: Sciences Biology *Homology (biology), any characteristic of biological organisms that is derived from a common ancestor *Sequence homology, biological homology between DNA, RNA, or protein sequences * Homologous chrom ...
with mandibles in crustaceans and insects, although more recent studies (e.g. using Hox genes) suggest they are probably homologous with the crustacean second antennae.


Chelate pedipalps

Chelate or sub-chelate (pincer-like) pedipalps are found in several arachnid groups ( Ricinulei, Thelyphonida, scorpions and pseudoscorpions) but the chelae in most of these groups may not be homologous with those found in Xiphosura. The pedipalps are distinctly raptorial (i.e., modified for seizing prey) in the Amblypygi, Thelyphonida, Schizomida, and some Opiliones belonging to the laniatorid group.


Spider pedipalps

Pedipalps of
spiders Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species dive ...
have the same segmentation as the legs, but the tarsus is undivided, and the pretarsus has no lateral claws. Pedipalps contain sensitive chemical detectors and function as taste and smell organs, supplementing those on the legs. In sexually mature male spiders, the final segment of the pedipalp, the tarsus, develops a complicated structure (sometimes called the palpal bulb or palpal organ) that is used to transfer sperm to the female seminal receptacles during mating. The details of this structure vary considerably between different groups of spiders and are useful for identifying species. The pedipalps are also used by male spiders in courtship displays, contributing to vibratory patterns in web-shaking, acoustic signals, or visual displays. The cymbium is a spoon-shaped structure located at the end of the spider pedipalp that supports the palpal organ. The cymbium may also be used as a
stridulatory organ Stridulation is the act of producing sound by rubbing together certain body parts. This behavior is mostly associated with insects, but other animals are known to do this as well, such as a number of species of fish, snakes and spiders. The mech ...
in spider courtship."Seismic signal production in a wolf spider: parallel versus serial multi-component signals"
''Journal of Experimental Biology''.
The embolus is a narrow whip-like or leaf-like extension of the palpal bulb.


References


Further reading

* * * compiled by Stephen W. Nichols; including Supplement A by George S. Tulloch. Published by The New York Entomological Society in cooperation with the American Museum of Natural History.


External links

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- ''Brachypelma vagans'' (Mexican red-rump) * Phrynus longipes#Pedipalps {{Spider nav Arachnid anatomy Sex organs Spider anatomy